Raveniola hirta, Zonstein, 2024
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publication ID |
https://doi.org/ 10.5852/ejt.2024.967.2699 |
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publication LSID |
lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738 |
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DOI |
https://doi.org/10.5281/zenodo.14034686 |
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persistent identifier |
https://treatment.plazi.org/id/A86FF08F-F3AD-4AB3-B399-7ACC509D3548 |
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taxon LSID |
lsid:zoobank.org:act:A86FF08F-F3AD-4AB3-B399-7ACC509D3548 |
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treatment provided by |
Plazi |
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scientific name |
Raveniola hirta |
| status |
sp. nov. |
Raveniola hirta sp. nov.
urn:lsid:zoobank.org:act:A86FF08F-F3AD-4AB3-B399-7ACC509D3548
Figs 8 View Figs 1–9 , 42 View Figs 36–44 , 67–68 View Figs 63–68 , 90 View Figs 82–90 , 117 View Figs 109–117 , 144 View Figs 136–147 , 173–174 View Figs 172–183 , 209 View Figs 202–210 , 236 View Figs 229–237 , 266 View Figs 265–273 , 296 View Figs 290–309 , 313–314 View Figs 310–318 , 357 View Figs 349–363 , 400–402 View Figs 400–408 , 504–508 View Figs 504–521 , 571–572 View Figs 565–574 , 651–658, 752
Diagnosis
Within the concolor species group of Raveniola , the holotype male of R. hirta sp. nov. is well distinguishable, because it possesses the shortest embolus, compared to other species ( Figs 400–402 View Figs 400–408 cf. Figs 385–399 View Figs 379–388 View Figs 389–399 , 403–428 View Figs 400–408 View Figs 409–417 View Figs 418–428 ); it also has long modified hairs on the tibia and metatarsus IV, as shown in Fig. 313 View Figs 310–318 (vs their presence in those species, where these hairs are known, only in females).Considering the last character, this new species shares with R. afghana sp. nov., R. alajensis sp. nov. and R. karategensis sp. nov. the presence of long modified hairs on the female tibia and metatarsus IV. Females of R. hirta can be distinguished from those of R. alajensis and R. karategensis by having a uniformly coloured (vs fairly ornamented) abdomen ( Figs 42 View Figs 36–44 , 67–68 View Figs 63–68 cf. Figs 38, 44 View Figs 36–44 , 65 View Figs 63–68 ), and from those of R. afghana sp. nov. in possessing much shorter spermathecae provided with clearly wider inner branches ( Figs 504–508 View Figs 504–521 cf. Figs 489–490 View Figs 487–503 ).
Etymology
The specific epithet is an adjective referring to a hirsute (Latin: ‘ hirt-us/-a/-um ’) leg IV in this species, densely covered with elongated fine hairs in females and to a lesser extent in males; the gender is feminine.
Material examined
Holotype
TAJIKISTAN • ♂; Darvaz Mts (northern slope), upper part of Zidadara Canyon , 2.5 km NNE of Haburabot Pass; 38°38.8′ N, 70°43.6′ E; 2900 m a.s.l.; 13 Jul. 1988; S. Zonstein leg.; SMNH. GoogleMaps
Paratypes (28 ♀♀)
TAJIKISTAN • 7 ♀♀; same collection data as for holotype; 2900–3300 m a.s.l.; SMNH GoogleMaps • 6 ♀♀; same locality as for holotype; 2850 m a.s.l.; 11 Jul. 2019; S. Zonstein and A. Hakimov leg.; SMNH GoogleMaps • 7 ♀♀; Darvaz Mts (top watershed zone), Haburabot Pass ; 38°37.229′ N, 70°43.135′ E; 3300 m a.s.l.; subalpine meadow-steppe; 26–27 Jul. 2023; A.A. Fomichev leg.; ISEA GoogleMaps • 6 ♀♀; same collection data as for preceding; ZMMU GoogleMaps • 2 ♀♀; Darvaz Mts (southern slope), Obiviskharvi Canyon ; 38°34′ N, 71°03′ E; 3000–3300 m a.s.l.; 14 Jul. 1988; S. Zonstein leg.; SMNH GoogleMaps .
Description
Male (holotype)
HABITUS. See Fig. 8. View Figs 1–9
MEASUREMENTS. TBL 17.40, CL 6.83, CW 6.27, LL 0.55, LW 1.00, SL 3.57, SW 2.97.
COLOUR. Carapace laterally and posteriorly, palpal femur, entire leg I and femora II–IV cherry red, chelicerae, most part of cephalic portion, thoracic fovea and radial grooves of carapace darker reddish brown; other segments of palp and legs II–IV dark yellowish orange; clypeus and eye tubercle blackish brown; sternum, labium, maxillae and ventral surface of abdomen including spinnerets yellowish brown; abdomen dorsally uniformly brown.
CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 90 View Figs 82–90 . Clypeus and eye group as in Fig. 144 View Figs 136–147 . Eye diameters and interdistances: AME 0.16(0.20), ALE 0.27, PLE 0.25, PME 0.16; AME–AME 0.17(0.13), ALE–AME 0.14(0.12), ALE–PLE 0.13, PLE–PME 0.06, PME–PME 0.55. Chelicerae with weak rastellum composed in each basal segment of 35–40 thickened spikes in front of fang base. Each cheliceral furrow with 8 promarginal teeth and 4–5 mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in Fig. 209 View Figs 202–210 . Maxillae with 30–35 cuspules each.
LEGS. Tibia and metatarsus I as in Figs 266 View Figs 265–273 , 296 View Figs 290–309 . Tibia and metatarsus IV covered with modified hairs ( Fig. 313 View Figs 310–318 ). Scopula: entire and distal on metatarsi I–II; entire on tarsi I–II; entire but mixed with setae on tarsus III; sparse and widely divided on tarsus IV. Trichobothria: 2 rows of 8–11each on tibiae, 16–19 on metatarsi, 12–17 on tarsi, 10–11 on cymbium. PTC I–IV with 7–8 teeth on each margin.
SPINATION. Metatarsus I and tarsi I–IV aspinose. Palp: femur d4, pd1, rd1; patella p2; tibia d6, p3, r3, v5; cymbium d4–5 normal + 10–15 small. Leg I: femur d4; patella p1; tibia p3, pv3, r2, rv2+ 2M. Leg II: femur d4, pd3(2), rd2; patella p3(2); tibia p3, v8(7); metatarsus p3, v6. Leg III: femur d4, pd3, rd3; patella p2(1), r1; tibia d3, p3, r3, v7; metatarsus p5, pd3, r3, v6. Leg IV: femur d4(3), pd3, rd3(2); patella p1, r1; tibia p3, r 3, v 8(7); metatarsus p3, pd3, r5, v8.
PALP. Tibia, cymbium and copulatory bulb as shown in Fig. 357 View Figs 349–363 . Embolus short with basal portion provided with low keel and with hooked apical part ( Figs 400–404 View Figs 400–408 ).
SPINNERETS. See Fig. 571 View Figs 565–574 . PMS: length 0.39, diameter 0.16. PLS: maximal diameter 0.67; length of basal, medial and apical segments 0.77, 0.42, 0.41; total length 1.60; apical segment triangular.
| Femur | Patella | Tibia | Metatarsus | Tarsus | Total | |
|---|---|---|---|---|---|---|
| Palp | 3.83 (3.69) | 1.85 (1.81) | 3.04 (2.38) | – | 1.08 (2.37) | 9.80 (10.25) |
| Leg I | 6.25 (5.32) | 3.43 (2.95) | 4.72 (3.62) | 4.97 (3.24) | 2.80 (2.29) | 22.17 (17.42) |
| Leg II | 5.76 (4.79) | 3.04 (2.58) | 4.10 (2.97) | 4.68 (2.99) | 2.64 (2.26) | 20.22 (15.59) |
| Leg III | 4.69 (3.99) | 2.41 (2.18) | 3.30 (2.48) | 4.41 (3.15) | 2.27 (1.96) | 17.08 (13.76) |
| Leg IV | 6.34 (5.75) | 3.11 (3.21) | 5.16 (4.49) | 6.83 (5.23) | 3.12 (2.97) | 24.56 (21.65) |
Female (paratype)
HABITUS. See Fig. 42. View Figs 36–44
MEASUREMENTS. TBL 16.40, CL 6.41, CW 5.56, LL 0.66, LW 1.30, SL 3.49, SW 3.10.
COLOUR. Carapace, palpal femur and femora I–IV dark brownish orange; other segments of palp and legs I–IV lighter brownish orange; chelicerae dark cherry red; eye tubercle with wide and partially fused blackish brown rings around eyes; sternum, labium, maxillae and ventral surface of abdomen including spinnerets yellowish brown; abdomen dorsally uniformly brown as in male.
CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 117 View Figs 109–117 . Clypeus and eye group as in Fig. 173 View Figs 172–183 . Eye diameters and interdistances: AME 0.15(0.21), ALE 0.31, PLE 0.25, PME 0.17; AME–AME 0.16(0.10), ALE–AME 0.13(0.10), ALE–PLE 0.14, PLE–PME 0.07, PME–PME 0.47. Chelicerae with weak rastellum composed of 25–30 thickened spikes in front of fang base. Each cheliceral furrow with 8 promarginal teeth and 3–4 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 236 View Figs 229–237 . Maxillae with 29–34 cuspules each.
LEGS. Tibia and metatarsus IV covered by dense modified hairs as in Fig. 314 View Figs 310–318 . Scopula: distal on metatarsi I–II; entire on palpal tarsus and tarsi I–II; widely divided by setae on tarsus III; vestigial on tarsus IV. Trichobothria: 2 rows of 9–11 each on tibiae, 16–18 on metatarsi, 14–15 on tarsi, 11 on palpal tarsus. Palpal claw with 4 promarginal teeth. PTC I–IV with 8–9 teeth on each margin.
SPINATION. Palpal femur with 3–4 dorsal bristles instead of spines, femora I–IV with one basodorsal spine and 3–4 median and/or apical bristles; palpal patella and patellae I, II and IV, and tarsi I–IV aspinose. Palp: femur pd1; tibia v7(6); tarsus v2. Leg I: femur pd2; tibia v4; metatarsus v5. Leg II: femur pd3; tibia v4(3); metatarsus v6. Leg III: femur pd2(1), rd3(2); patella p2, r1; tibia p2(1), r2, v6(5); metatarsus p3, pd2, rd3, v7. Leg IV: femur rd1; tibia v6; metatarsus r1, v9.
SPERMATHECAE. Each of paired spermathecae asymmetrical with relatively short and wide base carrying two diverging branches: massive trapezoidal inner branch and thin spindle-like outer one ( Fig. 508 View Figs 504–521 ).
SPINNERETS. See Fig. 572 View Figs 565–574 . PMS: length 0.58, diameter 0.28. PLS: length of basal, medial and apical segments 1.31, 0.67, 0.66; total length 2.64; apical segment triangular.
Variation
Carapace length in females (n=11) varies from 5.18 to 7.07. For female paratypes, the variations in their colouration, structure of the eye group and conformation of the spermathecae are shown in Figs 67–68 View Figs 63–68 , 174 View Figs 172–183 , 504–507 View Figs 504–521 .
Ecology
The species inhabits short-grass meadows and meadow-steppes in the subalpine and alpine zones. All spiders, including the holotype male, were collected from their relatively deep (of 35–45 cm depth) burrows. See Figs 651–658 View Figs 651–658 .
Distribution
Known from two highland localities in Darvaz Mts, Tajikistan ( Fig. 752 View Figs 751–760 ).
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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