Raveniola pallens, Zonstein, 2024

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 55-57

publication ID	https://doi.org/ 10.5852/ejt.2024.967.2699
publication LSID	lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738
DOI	https://doi.org/10.5281/zenodo.14034708
persistent identifier	https://treatment.plazi.org/id/4F2510F5-833E-42C4-9C5D-C17B434FFF9B
taxon LSID	lsid:zoobank.org:act:4F2510F5-833E-42C4-9C5D-C17B434FFF9B
treatment provided by	Plazi
scientific name	Raveniola pallens
status	sp. nov.

Treatment

Raveniola pallens sp. nov.

urn:lsid:zoobank.org:act:4F2510F5-833E-42C4-9C5D-C17B434FFF9B

Figs 17 View Figs 10–18 , 52 View Figs 45–53 , 99 View Figs 91–99 , 126 View Figs 118–126 , 154 View Figs 148–159 , 184 View Figs 184–195 , 218 View Figs 211–219 , 245 View Figs 238–246 , 276 View Figs 274–281 , 367 View Figs 364–378 , 434–435 View Figs 429–438 , 530–531 View Figs 522–536 , 593–594 View Figs 593–601 , 755

Diagnosis

Raveniola pallens sp. nov. differs from other members of the same species group in possessing a wider eye group with relatively smaller and noticeably broadly spaced anterior median eyes ( Figs 154 View Figs 148–159 , 184 View Figs 184–195 cf. 151–153, 155, 182–183, 185). Unlike males of R. diluta sp. nov. and R. fedotovi , possessing an embolic keel, the holotype male of R. pallens lacks this structure; it differs from males of R. zyuzini sp. nov. in having a noticeably lesser curved metatarsus I and by a sparser scopula on metatarsi and tarsi I–II ( Fig. 276 View Figs 274–281 cf. Fig. 277 View Figs 274–281 ). Females of R. pallens sp. nov. are distinguishable due to having a widely divided ventral scopula on the palpal tarsus (vs entire one in females of the related species), as well as owing to their low, mound-like and widely spaced spermathecal trunks (vs the higher cone-shaped ones in other species of the group; Figs 530–531 View Figs 522–536 cf. Figs 526–529, 532–533 View Figs 522–536 ).

Etymology

The specific epithet is a Latin adjective (of any gender) meaning ‘light’ or ‘pallid’ and referring to the pale ground color of the specimens belonging to this species.

Material examined

Holotype

UZBEKISTAN • ♂; Zeravshan Mts (southwestern slope) , foothills below Amankutan Pass; 39°13.5′ N, 66°53.5′ E; 1000–1300 m a.s.l.; 9 Apr. 1991; S. Zonstein leg.; SMNH. GoogleMaps

Paratype

UZBEKISTAN • 1 ♀; same collection data as for preceding; 900 m a.s.l.; 7 Apr. 1989; SMNH GoogleMaps .

Additional material (1 ♀ subad., 1 juv.)

UZBEKISTAN • 1 juv.; same collection data as for preceding; 850 m a.s.l.; 9 Apr. 1991; SMNH GoogleMaps • 1 ♀ subad.; northern foothills of Turkestan Mts , Beshbuloq Canyon , vicinity of Turkmen Village; 39°57′ N, 68°30′ E; 600–700 m a.s.l.; 27 Jun. 1980; A.B. Nenilin leg.; SMNH GoogleMaps .

Description

Male (holotype)

HABITUS. See Fig. 17. View Figs 10–18

MEASUREMENTS. TBL 9.05, CL 3.65, CW 2.99, LL 0.25, LW 0.63, SL 1.82, SW 1.57.

COLOUR. Carapace, palps and legs light brownish orange; eye tubercle blackish brown; chelicerae light reddish brown; sternum, labium and maxillae yellow; abdomen uniformly greyish white without distinct dorsal pattern.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 99 View Figs 91–99 . Clypeus and eye group as in Fig. 154 View Figs 148–159 . Eye diameters and interdistances: AME 0.10(0.14), ALE 0.19, PLE 0.12, PME 0.08; AME–AME 0.10(0.06), ALE–AME 0.05(0.03), ALE–PLE 0.03, PLE–PME 0.03, PME–PME 0.28. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 7 promarginal teeth and 7–8 mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in Fig. 218 View Figs 211–219 . Maxillae with 9–11 cuspules each.

LEGS. Tibia and metatarsus I as in Fig. 276 View Figs 274–281 . Scopula: entire and distal on metatarsi I–II; entire on tarsus I; narrowly divided with setae on tarsus II; widely divided on tarsus III; rudimentary on tarsus IV. Trichobothria: 2 rows of 8–9 each on tibiae, 13–14 on metatarsi, 10–12 on tarsi, 8 on cymbium. PTC I–IV with 6–8 teeth on each margin.

SPINATION. Patella I and tarsi I–IV aspinose. Palp: femur d3, pd1; tibia p3, r1, v5; cymbium d4+5 smaller. Leg I: femur d3, pd2; tibia p2, pv1, rv2+2M; metatarsus v1. Leg II: femur d3, pd2; patella p1; tibia p3, v7(6); metatarsus p1, v6. Leg III: femur d4, pd3, rd2; patella p2, r1; tibia d2, p2, r2, v7; metatarsus d3, p3, r3, v7. Leg IV: femur d4, pd3, rd2; patella p1, r1; tibia d2, p3, r3, v8(7); metatarsus d2, p4, r3, v8(7).

PALP. Tibia, cymbium and copulatory bulb as shown in Fig. 367 View Figs 364–378 . Embolus lacking keel, corkscrew-shaped, thin and gradually tapering to its tip ( Figs 434–435 View Figs 429–438 ).

SPINNERETS. See Fig 593 View Figs 593–601 . PLS: maximal diameter 0.31; length of basal, medial and apical segments 0.49, 0.30, 0.27; total length 1.06; apical segment triangular.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	1.93 (2.53)	1.01 (1.43)	1.53 (1.86)	–	0.64 (1.54)	5.11 (7.36)
Leg I	3.18 (3.54)	1.89 (2.17)	2.58 (2.74)	2.57 (2.07)	1.63 (1.49)	11.85 (12.01)
Leg II	2.79 (3.38)	1.62 (1.90)	2.30 (2.28)	2.39 (2.14)	1.58 (1.47)	10.68 (11.17)
Leg III	2.54 (2.97)	1.31 (1.68)	1.84 (2.15)	2.51 (2.53)	1.54 (1.54)	9.74 (10.87)
Leg IV	3.15 (3.99)	1.64 (1.97)	2.67 (3.23)	3.56 (3.94)	1.83 (1.76)	12.85 (14.89)

Female (paratype)

HABITUS. See Fig. 52. View Figs 45–53

MEASUREMENTS. TBL 13.20, CL 5.07, CW 4.43, LL 0.43, LW 1.04, SL 2.62, SW 2.31.

COLOUR. As in male, with body even paler; legs I–IV uniformly very light yellowish tan and noticeably paler than yellowish orange carapace.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 126 View Figs 118–126 . Clypeus and eye group as in Fig. 184 View Figs 184–195 . Eye diameters and interdistances: AME 0.12(0.17), ALE 0.23, PLE 0.12, PME 0.06; AME–AME 0.23(0.18), ALE–AME 0.10(0.08), ALE–PLE 0.03, PLE–PME 0.03, PME–PME 0.63. Cheliceral rastellum absent. Each cheliceral furrow with 9 promarginal teeth and 2–3 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 245 View Figs 238–246 . Maxillae with 10–11 pointed (not rounded or blunt) cuspules each.

LEGS. Scopula: entire and distal on metatarsi I–II; narrowly divided on tarsus I; widely divided on palpal tarsus and tarsus II; absent on tarsi III–IV. Trichobothria: 2 rows of 9–10 each on tibiae, 14–15 on metatarsi, 12–14 on tarsi, 9 on palpal tarsus. Palpal claw with 3 promarginal teeth. PTC I–II and III–IV with 5 and 4–5 teeth on each margin, respectively.

SPINATION. Femora I–IV dorsally along midline with 2–3 spines and 2–3 bristles; palpal patella, patella I, and tarsi I–IV aspinose. Palp: femur d4, pd1; tibia p4(2), v8(7); tarsus v2. Leg I: femur pd1; tibia p2, v6; metatarsus v7(4). Leg II: femur pd1; patella p1; tibia p2(1), v6(5); metatarsus v6. Leg III: femur pd3, rd2; patella p2, r1; tibia d1, p2, r2, v7; metatarsus d5, p3, r2, v7. Leg IV: femur pd2, rd3(2); patella p1, r1; tibia d1, p2(3), r3(2), v7; metatarsus d2, p2, r3, v7.

SPERMATHECAE. Each of paired spermathecae small and low mound-like, with short and wide trunk that carries small and short outer branch diverging from spermathecal trunk close to its apex ( Figs 530–531 View Figs 522–536 ).

SPINNERETS. See Fig. 594 View Figs 593–601 . PLS: maximal diameter 0.51; length of basal, medial and apical segments 0.91, 0.66, 0.42; total length 1.99; apical segment triangular.

Ecology

Raveniola pallens sp. nov. inhabits rocky slopes in foothills and the low-mountain zone between 600 and 1300 m a.s.l., covered by steppe grasslands and sparse shrubland with Pistacea vera L., Cercis griffithii , species of Acer , Prunus (sect. Amygdalus (L.) Benth. & Hook.f.), etc. By their nature, these biotopes resemble the landscape which is shown in Fig. 707 View Figs 707–714 .All spiders were found hiding under stones.

Distribution

Southeastern Uzbekistan. See Fig. 755 View Figs 751–760 .