Raveniola kirgizica, Zonstein, 2024

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 64-66

publication ID	https://doi.org/ 10.5852/ejt.2024.967.2699
publication LSID	lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738
DOI	https://doi.org/10.5281/zenodo.14034716
persistent identifier	https://treatment.plazi.org/id/11E652E9-0591-46EB-AEBD-97E713558C10
taxon LSID	lsid:zoobank.org:act:11E652E9-0591-46EB-AEBD-97E713558C10
treatment provided by	Plazi
scientific name	Raveniola kirgizica
status	sp. nov.

Treatment

Raveniola kirgizica sp. nov.

urn:lsid:zoobank.org:act:11E652E9-0591-46EB-AEBD-97E713558C10

Figs 20 View Figs 19–27 , 55 View Figs 54–62 , 102 View Figs 100–108 , 129 View Figs 127–135 , 157 View Figs 148–159 , 187 View Figs 184–195 , 221 View Figs 220–228 , 248 View Figs 247–255 , 280 View Figs 274–281 , 304 View Figs 290–309 , 371 View Figs 364–378 , 441–442 View Figs 439–447 , 537 View Figs 537–554 , 601–603 View Figs 593–601 View Figs 602–610 , 717–718, 756

Diagnosis

In possessing a gently twisted embolus, the male holotype of R. kirgizica sp. nov. resembles males of R. kopetdaghensis , R. mikhailovi and R. virgata . It can be distinguished from the former species in the presence of PMS (vs their absence in R. kopetdaghensis ) and from the two latter species in possessing a noticeably longer tibia and metatarsus I ( Fig. 280 View Figs 274–281 cf. Figs 282–283, 286–287 View Figs 282–289 ); the detailed structure of the embolus also looks different ( Figs 441–442 View Figs 439–447 cf. Figs 443–446 View Figs 439–447 , 454–458 View Figs 448–456 View Figs 457–465 ). The only known female (paratype) is distinguishable due to a specific conformation of the paired spermathecae, which are so small and thin, and so widely spaced from each other, that they can be reliably distinguished from the spermathecae in all other species of this group ( Fig. 537 View Figs 537–554 cf. Figs 334–536 View Figs 334–348 View Figs 349–363 View Figs 364–378 View Figs 379–388 View Figs 389–399 View Figs 400–408 View Figs 409–417 View Figs 418–428 View Figs 429–438 View Figs 439–447 View Figs 448–456 View Figs 457–465 View Figs 466–474 View Figs 475–486 View Figs 487–503 View Figs 504–521 View Figs 522–536 , 538–554 View Figs 537–554 ).

Etymology

The specific epithet is a toponym referring to the range of this species: Kyrgyzstan (also Kirgizia).

Material examined

Holotype

KYRGYZSTAN • ♂; Alay Mts (northern slope), Kyrgyz-Ata Canyon , Karagoi Gorge ; 40°03′ N, 72°36′ E; 2450 m a.s.l.; 22 May 1993; S. Zonstein leg.; SMNH. GoogleMaps

Paratype

KYRGYZSTAN • 1 ♀; Alay Mts (northern slope), Beleuli Canyon , 15 km SW of Gulcha Town, environs of Chon-Beleuli Village; 40°14′ N, 73°37′ E; 2200 m a.s.l.; 28 Jul. 1988; S. Zonstein leg.; SMNH GoogleMaps .

Additional material

KYRGYZSTAN • 1 juv.; Alay Mts (northern slope), Kurshab-Gulcha Canyon , 1 km SSE of Gulcha Town, left bank of Gulcha River; 40°18′ N, 73°27′ E; 1800–2000 m a.s.l.; 11 Aug. 1985; S. Zonstein leg.; SMNH GoogleMaps .

Description

Male (holotype)

HABITUS. See Fig. 20. View Figs 19–27

MEASUREMENTS. TBL 12.50, CL 5.53, CW 4.64, LL 0.47, LW 0.92, SL 2.49, SW 2.38.

COLOUR. Carapace (except for slightly darker postocular band and radial groves), palps and legs light brownish orange; eye tubercle blackish brown; chelicerae light red; sternum, labium and maxillae pale yellowish orange; abdomen light greyish brown, dorsally with weakly developed medium sepia brown chevron-like pattern; book-lungs and spinnerets pale yellowish brown.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 102 View Figs 100–108 . Clypeus and eye group as in Fig. 157 View Figs 148–159 . Eye diameters and interdistances: AME 0.13(0.18), ALE 0.18, PLE 0.16, PME 0.13; AME–AME 0.14(0.09), ALE–AME 0.10(0.08), ALE–PLE 0.08, PLE–PME 0.05, PME–PME 0.35. Anterior cheliceral edge with unmodified setae; rastellum not developed. Intercheliceral tumescence indiscernible. Each cheliceral furrow with 10–11 promarginal teeth and 3 mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in Fig. 221 View Figs 220–228 . Maxillae with 12 cuspules each.

LEGS. Tibia and metatarsus I as in Figs 280 View Figs 274–281 , 304 View Figs 290–309 . Scopula: entire and distal on metatarsi I–II; entire on tarsus I; narrowly divided by setae on tarsus II; widely divided on tarsus III; vestigial on tarsus IV. Trichobothria: 2 rows of 9–10 each on tibiae, 16–18 on metatarsi, 12–15 on tarsi, 11 on cymbium. PTC I–IV with 6–7 teeth on each margin.

SPINATION. Palp: femur d3, pd2, rd1; patella pd1; tibia d2, p3, r1, v12–15; cymbium d4. Leg I: femur d4, pd3, rd3; patella p1; tibia p2, pv1, r1, rv2+ 2M. Leg II: femur d4, pd3; patella p1; tibia p3, v7; metatarsus v5. Leg III: femur d4, pd3, rd3; patella p2, r1; tibia d3, p2, r2, v7; metatarsus d3, p3, r4, v9. Leg IV: femur d4, pd3, rd3; patella p1, r1; tibia d2, p3, r3, v7; metatarsus d4, p3, r3, v9. Metatarsus I and tarsi I–IV aspinose.

PALP. Tibia, cymbium and copulatory bulb as shown in Fig. 371 View Figs 364–378 . Embolus long tapering, clearly Sshaped in lateral view, and curved outwards along its entire length ( Figs 441–442 View Figs 439–447 ).

SPINNERETS. See Figs 601–602 View Figs 593–601 View Figs 602–610 . PMS: length 0.34, diameter 0.12. PLS: maximal diameter 0.43; length of basal, medial and apical segments 0.73, 0.46, 0.51; total length 1.70; apical segment shortly digitiform.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	2.89 (2.33)	1.44 (1.59)	2.41 (2.02)	–	0.87 (1.79)	7.61 (7.73)
Leg I	4.61 (3.64)	2.50 (2.17)	3.79 (2.79)	4.07 (2.24)	2.11 (1.65)	17.08 (12.49)
Leg II	4.32 (3.41)	2.21 (2.09)	3.63 (2.51)	3.60 (2.29)	2.09 (1.68)	15.85 (11.98)
Leg III	4.07 (2.94)	1.71 (1.78)	3.22 (2.21)	4.24 (2.63)	2.34 (1.75)	15.58 (11.31)
Leg IV	5.03 (4.10)	2.18 (2.17)	4.25 (3.09)	5.96 (3.96)	2.63 (2.01)	20.05 (15.33)

Female (paratype)

HABITUS. See Fig. 55. View Figs 54–62

MEASUREMENTS. TBL 17.60, CL 5.32, CW 4.55, LL 0.53, LW 1.03, SL 2.48, SW 2.37.

COLOUR. Mostly as in male, but with noticeably paler dorsal abdomen. Dorsal pattern presented by only three pairs of short diffuse chevrons on posterior quarter of abdomen.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 129 View Figs 127–135 . Clypeus and eye group as in Fig. 187 View Figs 184–195 . Eye diameters and interdistances: AME 0.12(0.18), ALE 0.19, PLE 0.14, PME 0.12; AME–AME 0.17(0.11), ALE–AME 0.12(0.09), ALE–PLE 0.12, PLE–PME 0.05, PME–PME 0.43. Cheliceral rastellum absent. Each cheliceral furrow with 9 promarginal teeth and 2 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 248 View Figs 247–255 . Maxillae with 12–13 cuspules each.

LEGS. Scopula: entire and distal on metatarsi I–II; entire on palpal tarsus; narrowly divided with setae on tarsus I; widely divided on tarsus II; rudimentary and bilateral on tarsus III; absent on tarsus IV. Trichobothria: 2 rows of 8–9 each on tibiae, 13–15 on metatarsi, 12–14 on tarsi, 10 on palpal tarsus. Palpal claw with 3 promarginal teeth. PTC I–IV with 5–7 teeth on each margin.

SPINATION. Femora I–IV with one basodorsal spine and 4–5 dorsal spikes; palpal femur dorsally with 4 spikes; patellae I–II with 1 prodorsal spike; palpal patella, and tarsi I–IV aspinose. Palp: femur pd1; tibia p2, v8(7); tarsus v5(4). Leg I: femur pd1; tibia p2, v7; metatarsus p1(0), v7(6). Leg II: femur pd1; tibia p2, v7; metatarsus v7. Leg III: femur pd2, rd2; patella p2, r1; tibia d1, p2, r2, v7; metatarsus d4, p3, r3, v7. Leg IV: femur pd1, rd1; patella p1, r1; tibia d1, p2, r2, v7; metatarsus d3, p4, r4, v8.

SPERMATHECAE. Individual spermathecae widely spaced from one another. Each of paired spermathecae Y-shaped with relatively short and narrow base dividing into a pair of equally thin but unevenly long branches: shorter inner and longer outer ones ( Fig. 537 View Figs 537–554 ).

SPINNERETS. See Fig. 603 View Figs 602–610 . PMS: length 0.30, diameter 0.14. PLS: maximal diameter 0.53; length of basal, medial and apical segments 1.08, 0.62, 0.48; total length 2.18; apical segment triangular.

Ecology

The spiders were found under stones in sparse montane forest, dominated by Juniperus spp. (chiefly, by J. turkestanica Kom. and J. seravschanica ). The landscape and the natural biotope of the type locality is shown in Figs 717–718 View Figs 715–722 .

Distribution

Known only from the type locality. See Fig. 756 View Figs 751–760 .