Raveniola ignobilis, Zonstein, 2024

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 34-36

publication ID	https://doi.org/ 10.5852/ejt.2024.967.2699
publication LSID	lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738
DOI	https://doi.org/10.5281/zenodo.14026332
persistent identifier	https://treatment.plazi.org/id/17DD6055-088F-4BEF-A1D6-DD4686AF191C
taxon LSID	lsid:zoobank.org:act:17DD6055-088F-4BEF-A1D6-DD4686AF191C
treatment provided by	Plazi
scientific name	Raveniola ignobilis
status	sp. nov.

Treatment

Raveniola ignobilis sp. nov.

urn:lsid:zoobank.org:act:17DD6055-088F-4BEF-A1D6-DD4686AF191C

Figs 9 View Figs 1–9 , 43 View Figs 36–44 , 91 View Figs 91–99 , 118 View Figs 118–126 , 145 View Figs 136–147 , 175 View Figs 172–183 , 198 View Figs 196–201 , 210 View Figs 202–210 , 237 View Figs 229–237 , 267 View Figs 265–273 , 297 View Figs 290–309 , 315 View Figs 310–318 , 358–359 View Figs 349–363 , 403–408 View Figs 400–408 , 509 View Figs 504–521 , 573–574 View Figs 565–574 , 629– 631, 752

Diagnosis

Due to its relatively short copulatory bulb, males of Raveniola ignobilis sp. nov. resemble the holotype of R. dolosa sp. nov., but differ from the latter in possessing two raised opposite keels in the proximal part of the embolus ( Figs 403–408 View Figs 400–408 cf. Figs 397–399 View Figs 389–399 ). The conspecific females are distinguishable by a specific structure of their spermathecae, with long inner and outer branches set very close to each other ( Fig. 509 View Figs 504–521 ) vs differently arranged branches in other related species, where these branches are either shorter ( Figs 500–503 View Figs 487–503 , 524–525 View Figs 522–536 ) or broader spaced ( Figs 495–499 View Figs 487–503 , 520–523 View Figs 504–521 View Figs 522–536 ).

Etymology

The specific epithet is a Latin adjective meaning ‘obscure’, ‘inglorious’ and referring to a rather middling (or averaged) appearance of these congeners.

Material examined

Holotype

TAJIKISTAN • ♂; Hazratisho Mts , Kapar (Sangdara) Gorge, environs of Sangdara Village; 38°22′ N, 70°10′ E; 1650 m a.s.l.; 15 Oct. 1987; S. Zonstein leg.; SMNH. GoogleMaps

Paratypes (4 ♂♂, 1 ♀, 1 ♀ subad.)

TAJIKISTAN • 4 ♂♂, 1 ♀; same collection data as for preceding; 1450–1800 m a.s.l.; SMNH GoogleMaps • 1 ♀ subad.; same collection data as for preceding, Iokunj Gorge ; 38°23′ N, 70°09′ E; 1700 m a.s.l.; 18 May 2002; S. Zonstein leg.; SMNH GoogleMaps .

Additional material (1 ♀ subad., 1 juv.)

TAJIKISTAN • 1 ♀ subad.; Hazratisho Mts. , 25 km E of Muminabad Town; 24 May 1966; E.M. Andreeva leg.; MIZW • 1 juv.; Darvaz Mts , Hirakdara Canyon between Kalai-Husain and Safedoron Villages; 2300 m a.s.l.; Juniperus park forest; 4 Jul. 1970; E.M. Andreeva leg.; MIZW .

Description

Male (holotype)

HABITUS. See Fig. 9. View Figs 1–9

MEASUREMENTS. TBL 14.40, CL 5.96, CW 5.59, LL 0.45, LW 0.89, SL 3.16, SW 2.87.

COLOUR. Carapace and leg I from femur to basal metatarsus intensely brownish orange; other parts of leg I, and entire palps and legs II–IV, as well as sternum, labium and maxillae lighter brownish orange; eye tubercle with eyes surrounded with partially fused blackish rings, chelicerae light cherry red, abdomen pale greyish brown, dorsally with diffuse and weakly developed brownish chevron-like pattern; book-lungs and spinnerets very pale greyish yellow.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 91 View Figs 91–99 . Clypeus and eye group as in Fig. 145 View Figs 136–147 . Eye diameters and interdistances: AME 0.15(0.20), ALE 0.27, PLE 0.22, PME 0.16; AME–AME 0.14(0.09), ALE–AME 0.09(0.06), ALE–PLE 0.09, PLE–PME 0.04, PME–PME 0.32. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 9 promarginal teeth and 5–6 mesobasal denticles. Presumed intercheliceral tumescence poorly discernible, small, maculate and setose ( Fig. 198 View Figs 196–201 ). Sternum, labium and maxillae as shown in Fig. 210 View Figs 202–210 . Maxillae with ca 60 cuspules each.

LEGS. Tibia and metatarsus I as in Figs 267 View Figs 265–273 , 297 View Figs 290–309 . Scopula: entire and distal on metatarsi I–II; entire on tarsi I–II; widely divided on tarsi III–IV. Trichobothria: 2 rows of 9–10 each on tibiae, 13–15 on metatarsi, 12–13 on tarsi, 10 on cymbium. PTC I–II and III–IV with 8–9 and 10–12 teeth on each margin, respectively.

SPINATION. Palp: femur d4, pd1, rd1; patella pd1; tibia d4, p3, r1, v6; cymbium d3(5)+20–25 spikes. Leg I: femur d2+2 bristles, pd3; patella p1; tibia p2, pv2, rv2+ 2M. Leg II: femur d2+2 bristles, pd3; patella p1; tibia p3, v7; metatarsus p1, v5. Leg III: femur d4, pd3, rd2; patella p2(1), r1; tibia d4, p4, r3, v7; metatarsus d4, p4, r3, v6. Leg IV: femur d3, pd3, rd2; patella p2, r1; tibia d3, p4, r3, v7; metatarsus d1, p4, r3, v8. Tarsi I–IV aspinose.

PALP. Tibia, cymbium and copulatory bulb as shown in Figs 358–359 View Figs 349–363 . Embolus with long basal portion provided with low keel and short hooked apical part ( Figs 403–405 View Figs 400–408 ).

SPINNERETS. See Fig. 573 View Figs 565–574 . PMS: length 0.48, diameter 0.14. PLS: maximal diameter 0.51; length of basal, medial and apical segments 0.80, 0.49, 0.37; total length 1.66; apical segment triangular.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	3.74 (3.74)	1.89 (2.02)	2.92 (2.42)	–	0.97 (2.11)	9.52 (10.29)
Leg I	6.09 (4.84)	3.13 (2.96)	4.80 (3.52)	4.65 (2.64)	2.59 (1.77)	21.26 (15.73)
Leg II	5.70 (4.41)	2.81 (2.59)	4.58 (3.18)	4.39 (2.72)	2.55 (1.78)	20.03 (14.68)
Leg III	4.97 (3.72)	2.09 (2.01)	3.53 (2.45)	4.36 (3.17)	2.57 (1.93)	17.52 (13.28)
Leg IV	6.03 (4.87)	2.49 (2.62)	4.79 (3.51)	6.52 (4.55)	3.01 (2.43)	22.84 (17.98)

Female (paratype)

HABITUS. See Fig. 43. View Figs 36–44

MEASUREMENTS. TBL 16.60, CL 6.14, CW 5.58, LL 0.69, LW 1.19, SL 3.17, SW 2.86.

COLOUR. Similar to that of male, but dorsal abdominal chevron-like pattern darker and better developed.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 118 View Figs 118–126 . Clypeus and eye group as in Fig. 175 View Figs 172–183 . Eye diameters and interdistances:AME 0.16(0.22), ALE 0.30, PLE 0.17, PME 0.16; AME–AME 0.11(0.05), ALE–AME 0.10(0.07), ALE–PLE 0.09, PLE–PME 0.03, PME–PME 0.43. Cheliceral rastellum absent. Each cheliceral furrow with 8–9 promarginal teeth and 5–6 mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 237 View Figs 229–237 . Maxillae with 56–57 cuspules each.

LEGS. Tibia and metatarsus IV as shown in Fig. 315 View Figs 310–318 . Scopula: distal on metatarsi I–II; entire on palpal tarsus and tarsi I–II; sparse and widely divided by setae on tarsus III; rudimentary on tarsus IV. Trichobothria: 2 rows of 7–8 each on tibiae, 13–15 on metatarsi, 12–14 on tarsi. Palpal claw with 4 promarginal teeth. PTC I–II and III–IV with 4–5/5–6 and 6–7/8–9 teeth on inner/outer margins, respectively.

SPINATION. All femora with one basodorsal spine and 3–4 median and/or apical bristles; palpal patella, patella I, and tarsi I–IV aspinose. Palp: femur pd1; tibia v7(6); tarsus v3(2). Leg I: femur pd2; tibia v3; metatarsus v6. Leg II: femur pd3; patella p1; tibia p2, v3; metatarsus v6. Leg III: femur pd3, rd3(2); patella p2, r1; tibia d1, p2, r2, v7; metatarsus d4, p3, r3, v7. Leg IV: femur pd1, rd1; patella p1, r1; tibia d1, p3, r3, v7; metatarsus d1, p4, r4, v8.

SPERMATHECAE. Each of paired spermathecae V-shaped with relatively short and wide base carrying two closely arosing, equally thick, long and weakly diverging branches ( Fig. 509 View Figs 504–521 ).

SPINNERETS. See Fig. 574 View Figs 565–574 . PMS: length 0.61, diameter 0.23. PLS: maximal diameter 0.63; length of basal, medial and apical segments 1.06, 0.56, 0.28; total length 1.90; apical segment triangular.

Variation

Carapace length in the paratype males (n =4) varies from 5.49 to 6.96. Some variations in the structure of the copulatory bulb as shown in Figs 406–408 View Figs 400–408 .

Ecology

The spiders were collected from their retreats under stones in different montane biotopes, mostly from the mixed woodlands at 1400–2300 m, dominated by Juglans regia and Juniperus seravschanica ( Figs 629– 631 View Figs 627–634 ).

Distribution

Tajikistan: Hazratisho and Darvaz Mts. See Fig. 752 View Figs 751–760 .