Raveniola ornatula, Zonstein, 2024

Zonstein, Sergei L., 2024, A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia, European Journal of Taxonomy 967, pp. 1-185 : 42-44

publication ID	https://doi.org/ 10.5852/ejt.2024.967.2699
publication LSID	lsid:zoobank.org:pub:C08B8027-50CC-417E-BCD4-5183B9FF6738
DOI	https://doi.org/10.5281/zenodo.14034696
persistent identifier	https://treatment.plazi.org/id/5B8115AD-5059-44AC-80CB-5B4CA05A8158
taxon LSID	lsid:zoobank.org:act:5B8115AD-5059-44AC-80CB-5B4CA05A8158
treatment provided by	Plazi
scientific name	Raveniola ornatula
status	sp. nov.

Treatment

Raveniola ornatula sp. nov.

urn:lsid:zoobank.org:act:5B8115AD-5059-44AC-80CB-5B4CA05A8158

Figs 12 View Figs 10–18 , 47 View Figs 45–53 , 94 View Figs 91–99 , 121 View Figs 118–126 , 148 View Figs 148–159 , 179 View Figs 172–183 , 213 View Figs 211–219 , 240 View Figs 238–246 , 270 View Figs 265–273 , 299–300 View Figs 290–309 , 362 View Figs 349–363 , 415–417 View Figs 409–417 , 517–519 View Figs 504–521 , 581–583 View Figs 575–583 , 674, 753

Diagnosis

The new species shares with R. ornata sp. nov. a rich dorsal abdominal pattern, a wide roundish sternum, small PMS and a more or less densely ornamented ventral surface of the abdomen, but can be distinguished from the latter in possessing an even denser ventral abdominal pattern ( Figs 12 View Figs 10–18 , 47 View Figs 45–53 , 213 View Figs 211–219 , 240 View Figs 238–246 , 581–583 View Figs 575–583 cf. Figs 11 View Figs 10–18 , 45–46 View Figs 45–53 , 212 View Figs 211–219 , 239 View Figs 238–246 , 577–580 View Figs 575–583 ). Males of R. ornatula sp. nov. differ from males of the latter species in having a shorter and wider basal section of the embolus ( Figs 415–417 View Figs 409–417 cf. Figs 412–414 View Figs 409–417 ). The conspecific females can be distinguished from females of R. ornata in possessing larger but less numerous maxillary cuspules, as well as shorter and stouter basal (inner) branches of the spermathecae ( Figs 240 View Figs 238–246 , 517–519 View Figs 504–521 cf. Figs 239 View Figs 238–246 , 513–516 View Figs 504–521 ).

Etymology

The specific epithet is a Latin adjective meaning ‘decorated’; the name refers to a very dense maculate dorsal and ventral abdominal pattern; it should also emphasize the similarity of this species to a closely related congener, Raveniola ornata sp. nov.

Material examined

Holotype

TAJIKISTAN • ♂; Hazratisho Mts , Yahsu Canyon , Sangdara Gorge; 38°22′ N, 70°10′ E; 1450 m a.s.l.; 15 Oct. 1987; S. Zonstein leg.; SMNH. GoogleMaps

Paratypes

TAJIKISTAN • 1 ♂, 5 ♀♀; same collection data as for preceding; 1450–1800 m a.s.l.; SMNH GoogleMaps .

Description

Male (holotype)

HABITUS. See Fig. 12. View Figs 10–18

MEASUREMENTS. TBL 11.35, CL 4.27, CW 3.94, LL 0.30, LW 0.81, SL 2.12, SW 2.01.

COLOUR. Carapace, palps and legs medium brownish orange (all appendages gradually lighten toward apices); eye tubercle with AMEs widely bordered and other eyes emarginated with blackish coloured cuticle, chelicerae reddish orange; sternum, labium and maxillae pale brownish yellow; abdomen yellowish brown with numerous brownish marks forming well-developed reticulate pattern on both dorsum and side; book-lungs and spinnerets pale yellowish brown.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 94 View Figs 91–99 . Clypeus and eye group as in Fig. 148 View Figs 148–159 . Eye diameters and interdistances: AME 0.14(0.20), ALE 0.26, PLE 0.14, PME 0.14; AME–AME 0.14(0.08), ALE–AME 0.09(0.06), ALE–PLE 0.10, PLE–PME 0.07, PME–PME 0.34. Anterior cheliceral edge with unmodified setae; rastellum not developed. Each cheliceral furrow with 7 promarginal teeth and 5–6 relatively large and raised mesobasal denticles. MIT indiscernible. Sternum, labium and maxillae as shown in Fig. 213 View Figs 211–219 . Maxillae with 30–31 cuspules each.

LEGS. Tibia and metatarsus I as in Figs 270 View Figs 265–273 , 299 View Figs 290–309 . Scopula: entire and distal on metatarsi I–II; entire on tarsus I; narrowly divided with setae on tarsus II; absent on tarsi III–IV. Trichobothria: 2 rows of 8–9 each on tibiae, 11–13 on metatarsi, 11–12 on tarsi, 7–8 on cymbium. PTC I–II and III–IV with 8–10 and 9–11 teeth on each margin, respectively.

SPINATION. Palp: femur d3, pd2; patella pd1; tibia d2, p3, r2, v6; cymbium d6(4). Leg I: femur d4, pd3, rd3(2); patella p1; tibia p2, pv2, rv2+2M; metatarsus v1(0). Leg II: femur d4, pd3; patella p1; tibia p3, v7(6); metatarsus v5(4). Leg III: femur d4, pd3, rd3(2); patella p2, r1; tibia d2, p3(2), r4(3), v7; metatarsus d2, p3, r3, v7. Leg IV: femur d4, pd3, rd2; patella p1, r1; tibia d2, p3, r3, v7; metatarsus d3, p4, r4, v9. Tarsi I–IV aspinose.

PALP. Tibia, cymbium and copulatory bulb as shown in Fig. 362 View Figs 349–363 . Embolus with short and conical basal portion lacking keel, and with even shorter and twisted apical part ( Figs 415–417 View Figs 409–417 ).

SPINNERETS. See Fig. 581 View Figs 575–583 . PMS: length 0.31, diameter 0.10. PLS: maximal diameter 0.38; length of basal, medial and apical segments 0.58, 0.43, 0.42; total length 1.43; apical segment shortly digitiform.

	Femur	Patella	Tibia	Metatarsus	Tarsus	Total
Palp	2.60 (2.41)	1.44 (1.40)	1.99 (2.02)	–	0.70 (1.43)	6.73 (7.26)
Leg I	4.28 (3.32)	2.19 (2.16)	3.14 (2.33)	3.60 (1.98)	2.15 (1.35)	15.36 (11.14)
Leg II	3.85 (3.07)	2.13 (1.98)	2.89 (2.07)	3.29 (1.96)	2.11 (1.33)	14.27 (10.41)
Leg III	3.23 (2.67)	1.59 (1.82)	2.25 (1.70)	3.34 (2.24)	2.08 (1.49)	12.49 (9.92)
Leg IV	4.37 (3.44)	1.91 (1.99)	3.16 (2.58)	4.65 (3.15)	2.33 (1.74)	16.42 (12.90)

Female (paratype)

HABITUS. See Fig. 47. View Figs 45–53

MEASUREMENTS. TBL 12.50, CL 4.32, CW 3.90, LL 0.37, LW 1.04, SL 2.08, SW 2.07.

COLOUR. As in male, but carapace and legs paler yellowish brown.

CEPHALOTHORAX. Carapace and chelicerae as shown in Fig. 121 View Figs 118–126 . Clypeus and eye group as in Fig. 179 View Figs 172–183 . Eye diameters and interdistances: AME 0.14(0.20), ALE 0.27, PLE 0.16, PME 0.12;AME–AME 0.16(0.10), ALE–AME 0.09(0.06), ALE–PLE 0.08, PLE–PME 0.08, PME–PME 0.32. Cheliceral rastellum absent. Each cheliceral furrow with 7 long promarginal teeth and 4–6 relatively large mesobasal denticles. Sternum, labium and maxillae as shown in Fig. 240 View Figs 238–246 . Maxillae with ca 35 cuspules each.

LEGS. Scopula: entire and distal on metatarsi I–II; narrowly divided on palpal tarsus; widely divided on tarsi I–II; absent on tarsi III–IV. Trichobothria: 2 rows of 7–8 each on tibiae, 8–10 on metatarsi, 10–11 on tarsi I–IV, 8 on palpal tarsus. Palpal claw with 6–7 promarginal teeth. PTC I–II and III–IV with 6–7 and 8–9 short teeth on each margin, respectively.

SPINATION. Palpal femur and femora I–II with 1 basodorsal spine and 3–4 dorsal bristles; palpal patella, patella I, and tarsi I–IV aspinose. Palp: femur pd1; tibia v7; tarsus v2. Leg I: femur pd1; tibia p1, v3; metatarsus v6(4). Leg II: femur pd1; patella p1; tibia p1, v5(4); metatarsus v7. Leg III: femur d4, pd3, rd3(2); patella p2, r1; tibia d1, p2, r1, v7; metatarsus p3, r3, v7. Leg IV: femur d4, rd1; patella p1(0), r1; tibia p2, r3, v7; metatarsus p3, r2, v7.

SPERMATHECAE. Each of paired spermathecae Y-shaped with relatively short and wide basic (inner) branch and diverging from this structure long outer branch ( Fig. 517 View Figs 504–521 ).

SPINNERETS. See Figs 582–583 View Figs 575–583 . PMS: length 0.32, diameter 0.09. PLS: maximal diameter 0.43; length of basal, medial and apical segments 0.85, 0.39, 0.38; total length 1.62; apical segment short digitiform.

Variation

Carapace length in the only male paratype is 4.18, in the female paratypes (n=5) it ranges from 3.98 to 4.83. Variation in the structure of the metatarsus I in male and the spermathecae as shown in Figs 300 View Figs 290–309 , 518–519 View Figs 504–521 .

Ecology

Raveniola ornatula sp. nov. was found in the midland mountain zone of Hazretisho Mts. where it inhabits (sympatrically with R. cucullata sp. nov. and R. ignobilis sp. nov.) shrubs and fragmentary broad-leaved woodlands dominated by Acer spp. and Juglans regia ( Fig. 674 View Figs 667–674 ). All spiders were found under stones.

Distribution

Known only from the type locality. See Fig. 753 View Figs 751–760 .