Anthemis rigida subsp. liguliflora (Halácsy) Greuter (1968: 263)

Goula, Katerina & Constantinidis, Theophanis, 2021, Taxonomic diversity and karyology of Anthemis rigida (Anthemideae, Asteraceae) in the Aegean, Greece, Phytotaxa 484 (1), pp. 129-143 : 140

Anthemis rigida subsp. liguliflora (Halácsy) Greuter (1968: 263) ( Fig. 5 : c,d)

Type:― GREECE. Cerigo [Kithira]: Hafen von Hagios-Nicolaos, 6 May 1902, Sterneck 279 ( WU 035428!) .

Annual with prostrate to ascending, simple or sparingly branched stems, 1.5−15 cm long. Stems and leaves greyishgreen, densely tomentose with medifixed hairs when young, less tomentose in fruit. Lower leaves long-petiolate, 1-pinnatisect, with narrowly pinnatifid, glandular-punctate segments; ultimate lobes obovate, usually twice as long as wide, subacute to shortly mucronate; upper leaves similar but smaller. Peduncles thickened, becoming rigid and usually recurved in fruit. Capitula solitary, radiate. Involucre 4−6(−7) × 3−4 mm, obconical to subcylindrical, bracts usually in 2−3 rows, imbricate, subequal in length, densely tomentose when young, thick, rigid, and often tinged purple in fruit. Outer involucral bracts narrowly lanceolate, 2.5−3.6 × ca. 1 mm, with scarious margins up to 0.2 mm. Inner bracts obovate, 2.7−3.8 × 1−1.2(−1.5) mm, with scarious margins (0.2−) 0.3−0.5 mm. Ligulate florets with ligules oblong-ovate, toothed at apex, 3.2−5.2 × 1−1.6 mm, white, glandular punctate, sterile; disc florets 2.5−3 mm, white, glandular punctate. Anthers, style, and pollen yellow. Receptacular scales hyaline, obovate to oblanceolate, mucronate and becoming lacerate at apex. Achenes obconical, faintly ribbed; outer achenes ca. 2 × 1 mm, with an auricle 0.5−0.9 mm long; inner achenes longer and thinner, ca. 2(−2.4) × 0.6−0.9 mm, with a corona 0.2−0.6 mm long.

Distribution and habitat:―This subspecies is endemic to the island of Kithira, where it grows on coastal habitats and stony places in phrygana communities.

Figures

FIGURE 5. Photographs of Anthemis rigida in the field (a, c, e, g) and in cultivation (b, d, f, h): A. rigida subsp. rigida from Kriti (a & b); A. rigida subsp. liguliflora from Kithira (c & d); A. rigida subsp. runemarkii from Milos (e & f); A. rigida subsp. antri-neptuni from Karpathos (g & h) (all photos taken by the first author except c, by Stavroula Fatsea).