Heteromeringia pectinata, Sasakawa, Mitsuhiro, 2011

Sasakawa, Mitsuhiro, 2011, Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae), Zootaxa 3038, pp. 1-28 : 22-23

Heteromeringia pectinata sp. nov.

( Figs 36–38 )

Type material. Holotype male ( BPBM 17251), Malaysia, Sabah, Ranau, 30. IX.–5. X. 1958, L. W. Quate. Paratypes: 3, same data as holotype; 3,Ƥ, same locality as holotype, 28–30. IX. 1958, T. C. Maa and 8–18. X. 1958, L. W. Quate; Ƥ, Sandakan, 20. XI. 1958, Maa; 3, 19 km N of Kalabakan, forest camp, 6. XI. 1962, Y. Hirashima, Malaise trap; 3, 9.8 km SW of Tenom, forest camp, 21. XII. 1962, K. J. Kuncheria;Ƥ, Malaysia, Sarawak, Kapit Distr., Merirai Valley, 28–31. VII. 1958, T. C. Maa.

Diagnosis. Largely black; face and gena brown; first antennal flagellomere largely brown; palpus entirely brown, with yellowish tip; anepisternum with yellow spot; dc three; surstylus with pectinate spines.

Description. MALE. Head with frontalia, ocellar triangle, occiput, and postgena shining black; parafrontalia, parafacialia, face, and gena brown (rarely yellow to yellowish brown); gena slightly whitish tomentose. Antenna brownish yellow, first flagellomere brown on apical 1/3 of outer side and almost entirely brown on inner side except for dorsoproximal corner (rarely entirely yellow), arista pale brown; palpus brown (slightly yellowish on apex) or entirely yellow. Thorax black, mesoscutum weakly shining, pleura and mediotergite strongly shining; postpronotal lobe brown, paler at anterior and posterior corners; katatergite brown. Wing hyaline, distinctly infuscated apically beyond middle of cell r1, but fading posteriorly beyond M1; calypter yellow, with fringe pale brown; halter yellow. Legs yellow but proximal 1/2–2/3 of mid coxa, distal 1/2–1/3 of fore femur, fore tibia, and fore tarsomeres 1–4 entirely brown to black. Abdomen with T1 yellowish brown, T2–epandrium shining black. Bristles dark brown to black.

Frons almost as wide as eye, slightly diverging ventrally to level of second or but slightly converging beyond that level; all or subequal in length, second and third or closely situated (distance between them less than 1/4 of distance between first and second or); oc short; poc long, more than 1/2 as long as ivt; frontalia with sparse minute setulae lateroventrally; gena 1/6–1/7 as high as eye; pm five to seven. First antennal flagellomere orbicular; arista sparsely but distinctly pubescent except for basal 1/5–1/6.

Mesoscutum with three dc, first dc about twice as long as acr and closely situated before second dc, second dc slightly shorter than third; eight to 10 rows of acr before transverse suture but just six or seven rows behind level of second dc; prsc absent; scutellum with bsc very short (1/8 length of asc). Wing 2.8–3.4 (3.4) mm long; three costal sections in proportion of 35: 10: 4.5, r-m at distal 1/2 of cell dm, M1 ratio 6.0, ultimate section of CuA1 5/6 as long as penultimate.

Epandrium ( Fig. 36 ) with long setae; surstylus swollen anterolaterally, with 14–16 pectinate spines in two rows; cerci united with each other; hypandrium with several setae; pre- and postgonites ( Fig. 37 ) each with one spine-like seta on tip. Epiphallus ( Fig. 38 ) weakly chitinized, mesophallus 950 µm long, hypophallus short and Vshaped, paraphallus with serration of seven minute teeth on distal sclerite, distiphallus J-shaped in lateral view; ejaculatory apodeme 300–320 µm long, 160–180 µm broad.

Body length 3.5–4.0 (4.0) mm.

FEMALE. Similar to male, but antennal scape and pedicel pale brown, palpus distinctly yellow apically, fore femur brown on distal 1/4 or faintly brown-tinged distally, fore tarsomere 4 yellow, T4–6 black, T7 brown, ovipositor brownish yellow; wing length 3.0– 3.4 mm, body length 3.9–4.7 mm.

Distribution. Malaysia (Sabah, Sarawak).

Remarks. This species has a large, yellow, circular disc on the anepisternum (distinctly larger than those of other species), and distinctive long outer setae and distal pectinate spines on the surstylus.

Etymology. The specific name refers to the pectinate (Latin: pectinatus) spines on the surstylus.

Figures

FIGURES 36 – 41. Male genitalia of Heteromeringia pectinata sp. nov. (36 – 38, paratype) and H. spinulifera sp. nov. (39 – 41, paratype). 37, pre- and postgonite. Scale = 0.1 mm.