Pustulosia hornensis ( Schaffer, 1912 ) Harzhauser & Guzhov & Landau, 2023

Harzhauser, Mathias, Guzhov, Aleksandr & Landau, Bernard, 2023, A revision and nomenclator of the Cainozoic mudwhelks (Mollusca: Caenogastropoda: Batillariidae, Potamididae) of the Paratethys Sea (Europe, Asia), Zootaxa 5272 (1), pp. 1-241 : 154-158

Pustulosia hornensis ( Schaffer, 1912) nov. comb.

Figs 31A–H

* Cerithium (Granulolabium) hornense — Schaffer 1912: 153, pl. 51, figs 48–50.

Pirenella hornensis (Schaffer) — Papp 1952: 111, pl. 2, figs 1–12.

P [otamides]. hornensis (Schff.) — Sieber 1958: 135.

Pirenella hornense ( Schaffer, 1912) —Ĥlzl 1958: 192.

Pirenella hornensis ( Schaffer, 1912) — Steininger et al. 1971: 378, pl. 5, figs 1–2.

Granulolabium haeutligenense n. sp. —Pfister & Wegm̧ller 2007: 90, pl. 8, figs 1–6.

Granulolabium haeutligenense —Pfister et al. 2011: 165.

non Pirenella hornensis ( Schaffer, 1912) — Mikuž 2003: 98, pl. 4, figs 1–2 [=? undescribed Potamididae ].

Type material. Lectotype designated herein: F/1146, SL: 17.8 mm, MD: 6.7 mm, Krahuletz Museum (Eggenburg, Austria), specimen illustrated in Schaffer (1912: pl. 51, fig. 49) ( Figs 31A 1 –A 2) . Paralectotype: F/1145, SL: 20.3 mm, MD: 7.3 mm, Krahuletz Museum (Eggenburg, Austria), specimen illustrated in Schaffer (1912: pl. 51, fig. 48) ( Figs 31B 1 –B 2) .

Type locality. Mold ( Austria) .

Stratigraphy. Early Miocene, Eggenburgian.

Illustrated material. NHMW 1999z0004/0001, SL: 13.8 mm, MD: 5.3 mm, Gauderndorf ( Austria), illustrated in Schaffer (1912: pl. 51, figs 33–34) ( Figs 31C 1 –C 2). NHMW 1934/0001/0062c, SL: 17.0 mm, MD: 6.6 mm, Mold ( Austria) ( Figs 31D 1 –D 2). NHMW 1934/0001/0062a, SL: 23.1 mm, MD: 7.5 mm, Mold ( Austria) ( Figs 31E 1 –E 2). NHMW 1934/0001/0062b, SL: 22.7 mm, MD: 8.1 mm, Mold ( Austria) ( Figs 31F 1 –F 2). NHMW 1934/0001/0062d, SL: 20.9 mm, MD: 7.6 mm, Mold ( Austria) ( Figs 31G 1 –G 2). NMBE 5015424 (=D1967), SL: 10.5 mm, MD: 5.0 mm, Naturhistorisches Museum Bern ( Switzerland), Häutlingen ( Switzerland), holotype of Granulolabium haeutligenense Pfister & Wegm ̧ller 2007, illustrated in Pfister & Wegm ̧ller 2007: 90, pl. 8, fig. 1 ( Fig. 31H) .

Studied material. 7 spec., NHMW 1934/0001/0062, Mold ( Austria) .

Revised description. Shell medium sized, moderately slender conical to weakly cyrtoconoid shell of about ten teleoconch whorls; apical angle ~35°, later decreasing to ~25°. Protoconch not preserved. Early teleoconch whorls with two prominent spiral cords with large, rounded beads. Third weaker spiral cord appears along abapical suture on third whorl, strengthening abapically and almost equal in strength on later whorls. Up to four delicate spiral threads between primary cords. Whorl profile subcylindrical; suture narrowly incised, undulating. Last whorl weakly convex, with convex, moderately constricted base bearing about four subobsoletely beaded spiral cords. Aperture subcircular-ovate; outer lip not thickened, without granules within; siphonal canal short, moderately wide, slightly deflected to the left; anal canal weakly incised. Columella excavated, smooth. Columellar callus forming broad rim, sharply delimited from base.

FIGURE 33. A 1 –A 2. Pustulosia submitralis ( Eichwald, 1851) , PM SPSU 3/318a, lectotype, ‘Volhynie’ probably Zalistsi (Ternopil Region, Ukraine), Middle Miocene, Badenian. B 1 –B 2. Pustulosia submitralis ( Eichwald, 1851) , NHMW (no number), Bernhardsthal ( Austria), Middle Miocene, Badenian. C 1 –C 2. Pustulosia submitralis ( Eichwald, 1851) , RGM 794074, Tilkikaya ( Turkey), Middle Miocene, Serravallian. D 1 –D 2. Pustulosia submitralis ( Eichwald, 1851) , NHMW 1859/0045/094, Oles'ko (Lviv Region, Ukraine), Middle Miocene, Badenian. E 1 –E 2. Pustulosia submitralis ( Eichwald, 1851) , NHMW 1862/0018/0320, Hidas ( Hungary), Middle Miocene, Badenian. F 1 –F 2. Pustulosia submitralis ( Eichwald, 1851) , NHMW 1859/0045/094, Oles'ko (Lviv Region, Ukraine), Middle Miocene, Badenian. G 1 –G 2. Pustulosia submitralis ( Eichwald, 1851) , NHMW 1859/0042/0026, Hidas ( Hungary), Middle Miocene, Badenian. H 1 –H 2. Pustulosia submitralis ( Eichwald, 1851) , NHMW 1859/0042/0026, Hidas ( Hungary), Middle Miocene, Badenian.

Synonyms. Granulolabium haeutligenense Pfister & Wegm ̧ller, 2007 is based on tectonically deformed specimens with two prominent spiral rows of beads accompanied by a weaker third row at the abapical suture. We interpret the plate-like morphology of the nodes, mentioned by Pfister & Wegm̧ller (2007) as characteristic for this species, as compressional deformation.

Discussion. Pustulosia hornensis is very close to Pustulosia submitralis ( Eichwald, 1851) and one might be inclined to synonymize both at first sight. Constant differences between them, however, are the presence of a prominent third spiral cord of beads at the abapical suture in P. hornensis , the prominent secondary spiral threads between the primaries, and its generally broader shape. In addition, they are separated by a stratigraphic gap of about 2 my. Nevertheless, both species might form a phylogenetic lineage.

Distribution. Pustulosia hornensis is endemic to the Central Paratethys Sea and is known so far only from the Eggenburgian (early Burdigalian) of the North Alpine Foreland Basin.

Central Paratethys. Eggenburgian (Early Miocene): North Alpine Foreland Basin: Häutlingen ( Switzerland) (Pfister et al. 2011); Kaltenbachgraben, Thalberggraben ( Germany) (Ĥlzl 1958); Mold, Gauderndorf ( Austria) ( Schaffer 1912).