Granulolabium moravicum (Hörnes, 1855)

Harzhauser, Mathias, Guzhov, Aleksandr & Landau, Bernard, 2023, A revision and nomenclator of the Cainozoic mudwhelks (Mollusca: Caenogastropoda: Batillariidae, Potamididae) of the Paratethys Sea (Europe, Asia), Zootaxa 5272 (1), pp. 1-241 : 128-131

Granulolabium moravicum (Hörnes, 1855)

Figs 25A–E

* Cerithium Moravicum Ĥrn. — Ĥrnes 1855: 402, pl. 42, fig. 7.

C [erithium]. Moravicum M. Ĥrn. var.— Vetters 1910: 159.

Pirenella moravica (Ĥrnes) —Davoli 1990: 50, pl. 4, figs 15a–b.

Granulolabium plicatum (Bruguière, 1792) —Harzhauser 2002: 73, pl. 1, figs 19–20 [non Granulolabium plicatum (Bruguière, 1792) ].

Granulolabium plicatum moravicum (Ĥrnes, 1856) — Harzhauser 2003: 194.

non Cerithium cf. moravicum Ĥrn. — Bajarunas 1910: 258. non Potamides moravicus M. Ĥrnes — Csepreghy-Meznerics 1950: 28, pl. 1, fig. 13 [= Tiarapirenella tabulata (Ĥrnes,

1856)]. non Pirenella moravica Ĥrnes, 1856 — Strausz 1955a: 47, 168, text-figs 2, 8 [= Tiarapirenella tabulata (Ĥrnes, 1856) ]. non Potamides (Pirenella) moravicus Ĥrnes, 1856 — Strausz 1966: 147, pl. 7, figs 23–29, pl. 8, figs 5, 10–11, pl. 10, figs 9–10

[= Tiarapirenella tabulata (Ĥrnes, 1856) ]. non Potamides moravica (Ĥrnes) — Kókay 1966: 41, pl. 4, fig. 6 [= Tiarapirenella tabulata (Ĥrnes, 1856) ]. non Pirenella moravica Hörnes, 1856 — Bałuk 2006: 197, pl. 8, fig. 5 [= Tiarapirenella tabulata (Ĥrnes, 1856) ].

Type material. Lectotype designated herein: NHMW 2022/0058/0001, SL: 27.9 mm, MD: 9.9 mm, illustrated in Ĥrnes (1855: pl. 42, fig. 7), Figs 25A 1 –A 2.

Type locality. Znojmo ( Czech Republic) .

Stratigraphy. Early Miocene, Karpatian.

Illustrated material. NHMW 2022/0059/0001, SL: 25.6 mm, MD: 8.7 mm, Helfens ( Austria), Figs 25B 1 –B 2. NHMW 2022/0059/0002, SL: 24.2 mm, MD: 8.3 mm, Helfens ( Austria), Figs 25C 1 –C 2. NHMW2000z0001/0057, SL: 29.9 mm, MD: 9.5 mm, Hipples ( Austria), illustrated in Harzhauser (2002: pl. 1, fig. 20), Figs 25D 1 –D 2. NHMW 2011/0010/0004, SL: 27.9 mm, MD: 9, 2 mm, Gebmanns ( Austria), illustrated in Sieber (1937: pl. 25, fig. E1), Figs 25E 1 –E 2. All Karpatian (Early Miocene).

Additional material. Karpatian (late Burdigalian): 33 spec., NHMW 2013/0293/0010, Kleinebersdorf ( Austria) ; 22 spec., NHMW 2013/0293/0004, Kleinebersdorf ( Austria) ; 24 spec., NHMW 2013/0293/0006, Kleinebersdorf ( Austria) ; 42 spec., NHMW 2022/0059/0003, Helfens ( Austria) ; 11 spec., NHMW 2022/0060/0001. Hipples ( Austria) ; 33 spec., NHMW 2022/0059/0004, Helfens ( Austria) .

Revised description. Shell medium sized, moderately slender, of about 12 teleoconch whorls, with weakly gradate spire; aperture angle ~30–35°. Protoconch and first teleoconch whorls unknown. Early spire whorls with two spiral rows of large, close-set, subquadratic beads. Third spiral cord at abapical suture usually fully covered by subsequent whorl. Secondary spiral cord of beads appears between primaries about 7 th or 8 th teleoconch whorls. Whorl profile subcylindrical. Suture slightly undulating, irregular. Whorl height slightly allometrically increasing during ontogeny, resulting in increasing number of spiral cords appearing at abapical suture on late spire whorls. Beads on adapical spiral cord usually more prominent, slightly pointed. Penultimate and last whorls with distinct primary and secondary spiral cords. Base moderately constricted, convex with indistinctly beaded, prominent primary and secondary spiral cords. Aperture relatively narrow ovate; outer lip not thickened, with spirally arranged granulae on inner aspect; anal canal narrowly incised; siphonal canal short, moderately wide, slightly deflected to the left. Columella excavated, smooth. Columellar callus forming broad rim.

Synonyms. None.

Discussion. The frequent presence of granulae inside the aperture ( Fig. 25A 1) supports the placement of this species in Granulolabium . This species seems to be an endemic Karpatian offshoot of Granulolabium plicatum and was variously treated as synonym or subspecies of G. plicatum by Harzhauser (2002, 2003), following the species concept of Lozouet (1986). The available material, however, differs constantly from G. plicatum . Granulolabium moravicum differs in its two dominant rows of beads on early teleoconch whorls, displays distinctly different primary and secondary spiral cords on the last whorl, is smaller and less slender than G. plicatum and has a relatively shorter spire. Especially the allometric increase of whorl height distinguished G. moravicum distinctly from G. plicatum . Plicatum -like morphs do not occur in the populations of G. moravicum . The paleogeographically restricted Paratethyan basins during the Karpatian might have favored the development of this endemic species similar to the development of endemic species in the Mainz Basin during the Oligocene as described by Kadolsky (1995).

Distribution. Cerithium moravicum Ĥrnes, 1855 is only documented from Karpatian (late Burdigalian) occurrences from the Korneuburg Basin and the adjacent North-Alpine Carpathian Foreland Basin.

Central Paratethys Sea. Karpatian (Early Miocene): Korneuburg Basin: Gebmanns, Helfens, Kleinebersdorf, Karnabrunn, Niederkreuzstetten, Obergänserndorf, Ŗckersdorf, Stetten, Wetzleinsdorf ( Austria) (Harzhauser 2002); North Alpine-Carpathian Foredeep: Znojmo ( Czech Republic) ( Ĥrnes 1855); Laa, Niederschleintz, Neuruppersdorf ( Austria) ( Vetters 1910; Sieber 1937; Papp 1952).