Mesohalina margaritacea ( Brocchi, 1814 ), 2001

Harzhauser, Mathias, Guzhov, Aleksandr & Landau, Bernard, 2023, A revision and nomenclator of the Cainozoic mudwhelks (Mollusca: Caenogastropoda: Batillariidae, Potamididae) of the Paratethys Sea (Europe, Asia), Zootaxa 5272 (1), pp. 1-241 : 113-118

Mesohalina margaritacea ( Brocchi, 1814)

Figs 20A–F

* Murex margaritaceus nob. — Brocchi 1814: 447, pl. 9, fig. 24. Cerithium lemniscatum — Andrzejowski 1832: 543 [non Brongniart, 1823]. Cerithium margaritaceum Brocc. — Pusch 1836: 525. Cerithium margaritaceum Brocc. — Pusch 1837: 148. C [erithium]. lemniscatum Brongn. — Pusch 1837: 149 [non Brongniart, 1823]. [ Cerithium ] margaritaceum Lam. — Ĥrnes 1848: 21. Cerithium margaritaceum Brocc. — Ĥrnes 1855: 403, pl. 42, figs 9a–b. Cerithium margaritaceum Brocc. — Fuchs 1868: 216. [ Potamides (Tympanotomus) margaritaceus Brocc. var. Nonndorfensis Sacc. — Sacco 1895: 46. Potamides margaritaceus Brocc. — Wolff 1897: 270, pl. 25, figs 18–21. Potamides submargaritaceus Braun. —B̂ckh 1899: 32, pl. 9, fig. 12a. Potamides margaritaceus Brocc. —B̂ckh 1899: 31, pl. 9, fig. 12b. Cerithium (Tympanotomus) margaritaceus Brocc. var. Nondorfensis [sic] Scc.— Schaffer 1912: 154, pl. 52, figs 1–2. Cerithium (Tympanotomus) margaritaceum Brocc. var. Nondorfensis [sic] Scc. — Schaffer 1913: 109, pl. 5, fig. 10. Cerithium margaritaceum — Petraschek 1926: 286, pl. 4, fig. 3. Cerithium (Tympanotomus) margaritaceum Brocc. — Munda 1939: 109, pl. 1, figs 16–17, pl. 2, figs 4–5. Potamides (Tympanotonus) calcaratus (Grateloup) Sacco — Stchepinsky 1939: 33, pl. 10, figs 12–15. Potamides (Tympanotonus) margaritaceus Brocchi — Stchepinsky 1941: 28, pl. 4, figs 14–16. Tympanotonus margaritaceus (Brocchi) — Stchepinsky 1946: 137, pl. 30, figs 3–6. Tympanotonus (Tympanotonus) margaritaceus (Brocchi) — Anić 1952: 40, pl. 10, figs 10–12. Tympanotonus (Tympanotonus) margaritaceus (Brocchi) 1814 — Rossi-Ronchetti 1955: 128, fig. 62. Potamides (Tympanotonos) margaritaceus Brocchi — Strausz 1956: 279, pl. 40, figs 16–24. Tympanotonus margaritaceus var. (?) submargaritaceus (Braun) — Seneš 1958: 138, pl. 21, fig. 280. Tympanotonus margaritaceus type; moniliformis — Seneš 1965: 69, text-fig. 1/3. Tympanotonus margaritaceus type; grateloupi — Seneš 1965: 69, text-fig. 1/4. Tympanotonus margaritaceus grateloupi (d’Orbigny 1852) —Ĥlzl 1958: 193, pl. 18, fig. 5. Tympanotonus margaritaceus calcarata — Seneš 1965: 69, text-fig. 1/2. Potamides margaritaceus Brocchi — Vadász 1960: 596, pl. 41, fig. 10. Cerithium margaritaceum Brocc. — Gamkrelidze et al. 1964: 231. Tympanotonus margaritaceus type — Seneš 1965: 69, text-fig. 1/1. Tympanotonus margaritaceum (Brocchi) 1824 —et forma calcarata (Grat.)—Ondrejičková & Seneš: 1965: 175, pl. 4, figs

42–45. Potamides (Tympanotonos) margaritaceus Brocchi — Strausz 1966: 163, pl. 78, figs 10–12, text-fig. 68. Tympanotonus (Tympanotonus) margaritaceus margaritaceus Brocchi — Moisescu 1969: 195, pl. 5, figs 67, 70–70 a, pl. 6, figs

72–77, 80. Tympanotonus (Tympanotonus) margaritaceus calcaratus (Grateloup in Sandberger) — Moisescu 1969: 198, pl. 5, figs 64–66. Tympanotonus margaritaceus margaritaceus Brocchi 1814 — Steininger et al. 1971: 379, pl. 5, figs 5–6. Tympanotonos (Tympanotonos) margaritaceus margaritaceus ( Brocchi, 1814) — Moisescu 1972: 93, pl. 30, figs 12a–b. Tympanotonos (Tympanotonos) margaritaceus calcaratum (Grateloup in Sandberger, 1863)— Moisescu 1972: 93, pl. 30, figs

13a–b. Tympanotonus (Tympanotonos) margaritaceus moniliforme (Grateloup in Sandberger, 1863)— Moisescu 1972: 93, pl. 30, figs

16–17. Tympanotonus margaritaceus ( Brocchi, 1814) — Báldi 1973: 260, pl. 29, figs 1–2. Tympanotonus margaritaceus nondorfensis [sic] Sacco 1895 — Steininger et al. 1973: 379, pl. 5, figs 4, 14–15. Tympanotonus margaritaceus moniliformis (Grateloup, 1840) — Steininger et al. 1973: 379, pl. 5, figs 7–9. Tympanotonus (Tympanotonus) margaritaceus (Brocchi) —Lebķchner 1974: 12, pl. 22, figs 4–5. Tympanotonus (Tympanotonus) margaritaceus var. nov.?—Lebķchner 1974: 12, pl. 22, figs 6–7. Tympanotonus margaritaceus Brocc. — Báldi & Steininger 1975: 344, pl. 10, fig. 1. Murex margaritaceus Brocchi, 1814 — Pinna & Spezia 1978: 151, pl. 37, fig. 3. Mesohalina margaritacea nondorfensis [sic] ( Sacco, 1895)— Wittibschlager 1983: 59, pl. 1, fig. 2. Mesohalina margaritacea calcarata (Grateloup, 1840) — Wittibschlager 1983: 59, pl. 1, figs 3a–b. Tympanotonus margaritaceus ( Brocchi, 1814) — Barthelt 1989: 39, pl. 12, fig. 1.

Tympanotonos margaritaceus grateloupi (d’Orbigny, 1852) — Moisescu 1992b: 19, pl. 3, figs 13a–b.

Mesohalina margaritacea margaritacea ( Brocchi, 1814) — Kadolsky 1995: 15, pl. 2, fig. 18.

Tympanotonos (T.) margaritaceus (Brocchi) — Taner 1996: 69, pl. 2, figs 3, 3a, 5, 5a.

Mesohalina („ Tympanotonus “) margaritacea nondorfensis [sic] Sacco— Schultz 1998: 54, pl. 20, fig. 16.

Tympanotonos margaritaceus grateloupi Orbigny — Popa & Chira 1999: 400, pl. 2, fig. 1.

Tympanotonos margaritaceus ( Brocchi, 1814) — Lozouet et al. 2001a: 26, pl. 9, figs 1–5.

Tympanotonos margaritaceus ( Brocchi, 1814) — Harzhauser & Kowalke 2001: 364, figs 5, 7–9.

Tympanotonos margaritaceus ( Brocchi, 1814) — Harzhauser & Mandic 2001: 696, pl. 1, figs 4–5.

Tympanotonos margaritaceus ( Brocchi, 1814) — Mikuž 2003: 101, pl. 4, figs 6–7.

Tympanotonos calcaratus (Grateloup, 1840) — Mikuž 2003: 103, pl. 4, figs 8–9.

Tympanotonos margaritaceus ( Brocchi, 1814) — Ýslamoðlu 2008: 273, figs 6/N–O.

Potamides (Mesohalina) margaritaceus ( Brocchi, 1814) — Esu & Girotti 2010: 154, pl. 6, figs 1–3.

Tympanotonos margaritaceus ( Brocchi, 1814) — Ýslamoðlu & Hakyemez 2010: 483, fig 7/e.

Tympanotonos margaritaceus (Brocchi) — Kovács & Vicián 2016: 246, pl. 1, fig. 6.

Mesohalina margaritacea ( Brocchi, 1814) — Harzhauser et al. 2016: 28, figs 11–13.

non Cerithium (Tympanotomus) margaritaceus Brocc. var. quadricincta Schff. — Schaffer 1912: 155, pl. 52, fig. 3 [= Terebralia

sp.]. non T [ympanotonus]. (T [ympanotonus].) margaritaceus quadricinctus Schff.— Sieber 1958: 136 [= Terebralia sp. ]. non Tympanotonus margaritaceus Brocchi, 1814 —Ģrsoy 2017: 79, pl. 1, figs 4a–c [= Ptychopotamides cinctus (Bruguière,

1792)].

Type material. Holotype, No. 5249, SL: 44.6 mm, MD: 20.9 mm, Museo di Storia Naturale Milano ( Italy).

Type locality. Unknown, maybe Mainz Basin (see above for genus).

Stratigraphy. Unknown, maybe Rupelian (see above for genus).

Illustrated material. NHMW 2022/0024/0003, SL: 58.9 mm, MD: 27.2 mm, Natural History Museum Vienna ( Austria), Máriahalom ( Hungary), Figs 20A. NHMW 1860/0050/0266, SL: 50.9 mm, MD: 24.2 mm, Natural History Museum Vienna ( Austria), Nonndorf ( Austria), Natural History Museum Vienna ( Austria), holotype of Potamides (Tympanotomus) margaritaceus nonndorfensis Sacco, 1895 , illustrated in Ĥrnes (1855: pl. 42, figs 9a–b), Figs 20B 1 –B 2. NHMW 1999z0004/0002, SL: 56.0 mm, MD: 26.0 mm, Natural History Museum Vienna ( Austria), Nonndorf ( Austria), Early Miocene, Eggenburgian, illustrated in Schaffer (1912, pl. 52, fig. 1), Figs 20C 1 –C 2. NHMW 2022/0192/0001, SL: 43.3 mm, MD: 22.8 mm, Kollmitzberg bei Adagger ( Austria), Egerian (late Oligocene), Fig. 20D. NHMW 2022/0192/0002, SL: 38.1 mm, MD: 26.6 mm, Kollmitzberg bei Adagger ( Austria), Egerian (late Oligocene), Fig. 20E. NHMW 2022/0193/0001, SL: 59.1 mm, MD: 29.9 mm, Máriahalom ( Hungary), Egerian (late Oligocene), Figs F 1 –F 2.

Studied material. Egerian (Chattian/Aquitanian): 58 spec., NHMW 2022/0024/0004, Máriahalom ( Hungary) ; Eggenburgian (Early Miocene): 21 spec., NHMW 1934/0001/0074, Maria Dreieichen ( Austria) ; 11 3 spec., NHMW 2022/0027/0001, Maria Dreieichen ( Austria) ; 11 spec., NHMW 1965/0000/0613, Mold ( Austria) ; 10 spec., NHMW 1845/0015/0049, Mold ( Austria) ; 7spec.,NHMW1846/0037/0375, Mold ( Austria) ; 8spec.,NHMW1866/0020/0001, Mold ( Austria) ; 5 spec., NHMW 2022/0028/0001, Nonndorf ( Austria) ; 8 spec., NHMW 1866/0040/0035, Nonndorf ( Austria) ; 10 spec., NHMW 2022/0029/0001, Nonndorf ( Austria) ; 8 spec., NHMW 2022/0030/0001, M̂rtersdorf ( Austria) ; 4 spec., NHMW 1860/0050/0371, Gauderndorf ( Austria) . Non-Paratethyan occurrences: Early Miocene: 7 spec., NHMW 2022/0031/0001, Bordeaux ( France) ; 6 spec. NHMW 1851/0017/1221, Dax ( France) ; 2 spec., NHMW 1883/0000/6081, La Brede ( France) ; 9 spec., NHMW 1867/0016/0002, Montpellier ( France) ; 8 spec., NHMW 1871/0010/0279; Oligocene : Alzey ( Germany) .

Revised description. Large conical shell of about 18 straight-sided teleoconch whorls with indistinct suture. Apical angle about 23–27°. Protoconch turreted, consisting of two strongly convex and smooth whorls of c. 200 μm diameter; second whorl with median keel. Onset of teleoconch marked by development of two spiral cords, separating whorl in steep subsutural ramp, weakly concave periphery and concave, strongly contracting abapical part. Suture narrow, sometimes accompanied by faint lower sutural thread. Orthocline to slightly orthocline-concave axial ribs cross spiral cords, forming regularly beaded sculpture and spirally elongate beads at intersections. Third spiral cord appears within 4 th –5 th teleoconch whorl below suture. Late teleoconch whorls with three prominent, densely beaded spiral cords, usually of more or less equal strength, but middle row weaker in some specimens. Egerian populations in Paratethys often with adapical row of wider spaced, blunt beads (calcarata morph). Two weaker secondary spiral rows of beads intercalated between primary cords on late teleoconch whorls; additional weak secondary row of beads at abapical suture, partly covered by subsequent whorl. One to three varices may appear on the last two whorls. Base convex, rapidly contracting covered by about six weakly beaded spiral cords. Aperture large, subquadratic-ovoid. Columella straight with central columellar fold of variable strength. Outer lip strongly thickened, callus, flaring, orthocline with protruding abapical margin of outer lip. Inner lip callus, abapically detached from base. Siphonal canal deeply incised, very narrow, recurved and strongly deflected to left. Anal canal forming prominent, broad, rounded notch adapically distorting outer lip, making it somewhat alate.

Discussion. Numerous subspecies and varieties of Mesohalina margaritacea have been described in the literature based on differences in sculpture (see Wittibschlager 1983). Herein, we follow Lozouet et al. (2001a), who treated many of these taxa as subjective junior synonyms of M. margaritacea . In Paratethyan material, the calcarata morph with prominent nodes along the adapical spiral cord is restricted to the Egerian. Harzhauser & Mandic (2001), however, documented co-occurring calcarata -morphs and typical M. margaritacea in the Egerian locality Kendl and observed prevailing calcarata morphs in the close-by, coeval locality Neuwinden ( Austria). Therefore, no geographic separation between these morphs can be stated. Oligocene populations from Máriahalom ( Hungary) do not differ is size and sculpture from Eggenburgian (early Burdigalian) populations from Nonndorf ( Austria). Therefore, no chrono-subspecies can be defined.

Cerithium (Tympanotomus) margaritaceus quadricincta Schaffer 1912 , from the Eggenburgian of Nonndorf ( Austria) is based on a fragmentary Terebralia specimen and must be excluded from M. margaritacea .

Distribution. Mesohalina margaritacea evolved during the Rupelian in the Proto-Mediterranean Sea. It is recorded from NE Italy (Castelgomberto; Menegatti 1978), the Tertiary Piedmont Basin in NW Italy (Carcare, Dego; Sacco 1888, 1895), the Denizli and Kale-Tavas basins in SW Turkey ( Ýslamoðlu 2008) and from the Thrace Basin ( Harzhauser et al. 2016). Rupelian occurrences in the Paris Basin and the North Sea Basin were separated by Marquet et al. (2008) as distinct subspecies M. margaritacea labyrinthum . Late Oligocene occurrences are reported from the NE Atlantic Aquitaine Basin ( Cossmann & Peyrot 1922) and throughout the Proto-Mediterranean Sea, e.g., southern Italy ( Esu & Girotti 2010), the Mesohellenic Basin in Greece ( Harzhauser 2004) and the Erzincan Basins in Turkey ( Stchepinsky 1941). At that time, the species extended also to the Mainz Basin ( Kadolsky 1995). In the Paratethys Sea, it appeared during the Chattian (Egerian) in the Bavarian and Austrian part of the North Alpine Foreland Basin ( Barthelt 1989, Harzhauser & Mandic 2002, Reichenbacher et al. 2004), in the Vértes Mountains and the Eger and Budapest regions in Hungary ( Seneš 1958, Báldi 1973, Janssen 1984, Leél-Őssy 1992) and in the Transylvanian Basin in Romania ( Moisescu 1972). It persisted up to the Eggenburgian (Early Burdigalian) in Austria, Slovakia and Slovenia ( Steininger et al. 1971, Mikuž 2003). During the Early Miocene, it was still widespread, occurring in the Aquitaine Basin ( Cossmann & Peyrot 1922, Lozouet et al. 2001a) in the Atlantic, the Rhône Basin in France ( Catzigras 1943), the Tertiary Piedmont Basin in Italy ( Sacco 1895), the Mesohellenic Basin in Greece ( Harzhauser & Kowalke 2001) and the Sivas Basin in Turkey ( Stchepinsky 1939, 1946). This species seems to have become extinct during the mid-Burdigalian.

Central Paratethys Sea. Egerian (late Oligocene/Early Miocene): North Alpine Foreland Basin: Hausham, Kaltenbachgraben, Thalberggraben ( Germany) (Ĥlzl 1962); Neuwinden, Zelking, Pielach, Viehdorf ( Austria) (Petraschek 1924, Harzhauser & Mandic 2001); Hungarian Paleogene Basin: Nagyegyháza, Nagymaros, Mány, Germely, Felsőörspuszta, Zsámbék, Tök, Szomor, Anyácsapuszta, Máriahalom, Vasztély, Csolnok, Tarján, Kesztölc, Solymár, Budafok, Szentendre, Pomáz, Leányfalu, Göd, Dömös, Diósjenő, Nagyoroszi, Borsosberény, Becska, Tolmács, Eger, Novaj, Mucsony, Sajókaza ( Hungary) (B̂ckh 1899; Báldi 1973); Esztergom Basin: Esztergom-Szentgyörgymező ( Hungary) ( Kovács & Vicián 2016); South Slovakian Basin: Štúrovo ( Slovakia) ( Ondrejíčková & Seneš 1965); Kováčov Basin: Kováčov ( Slovakia) ( Seneš 1958); Central Slovenia: Soteska ( Slovenia) ( Mikuž 2003); Transylvanian Basin: Zimbor ( Romania) ( Moisescu 1972). Eggenburgian (Early Miocene): Rhône Basin ( France) ( Mongin 1952); North Alpine-Carpathian Foreland Basin: Gauderndorf, Ķhnring, Maria Dreieichen, M̂rtersdorf, Mold, Nonndorf ( Austria) ( Steininger et al. 1973); Horná Nitra Basin: Veľká Čausa ( Slovakia) ( Steininger et al. 1973).

Proto-Mediterranean Sea–Central Paratethys junction: Burdigalian (Early Miocene): Rhône Basin ( France) ( Mongin 1952);

Eastern Paratethys. Middle Oligocene: Transcaucasia: Akhaltsikhe Depression ( Georgia) ( Gamkrelidze et al. 1964).

Early Miocene, Egerian. B 1 –B 2. Mesohalina margaritacea ( Brocchi, 1814) , NHMW 1860/0050/0266, Nonndorf ( Austria), Early Miocene, Eggenburgian, holotype of Potamides (Tympanotomus) margaritaceus nonndorfensis Sacco, 1895 . C 1 –C 2. Mesohalina margaritacea ( Brocchi, 1814) , NHMW 1999z0004/0002, Nonndorf ( Austria), Early Miocene, Eggenburgian. D.

Mesohalina margaritacea ( Brocchi, 1814) , NHMW 2022/0192/0001, SL: 43.3 mm, MD: 22.8 mm, Kollmitzberg bei Adagger

( Austria), Egerian (late Oligocene). E. Mesohalina margaritacea ( Brocchi, 1814) , NHMW 2022/0192/0002, SL: 38.1 mm, MD:

26.6 mm, Kollmitzberg bei Adagger ( Austria), Egerian (late Oligocene). F 1 –F 2. Mesohalina margaritacea ( Brocchi, 1814) , NHMW 2022/0193/0001, SL: 59.1 mm, MD: 29.9 mm, Máriahalom ( Hungary), Egerian (late Oligocene).

Unnamed batillariid family

In their molecular phylogeny of batillariid gastropods, Ozawa et al. (2009) showed that Batillariidae are restricted to the northwestern Pacific and Australasia. The neotropical Lampanella M̂rch, 1876 and Rhinocoryne Martens, 1900 , which have traditionally been placed in Batillariidae , are sister to the Planaxidae . Unfortunately, Ozawa et al. (2009) did not formally establish a family for this clade, which most probably would be named after Lampanella . As we do not have morphological and molecular data to describe this family in detail, we refrain from introducing a formal name.

Genus Lampanella Mörch, 1876

Type species. Murex minimus Gmelin, 1791 ; subsequent designation by Tryon (1887: 118). Recent, Western Atlantic.

Original diagnosis. “Taille petite; forme trapue: spire treillissée, à costules droits; dernier tour grand, arrondi à la base. Ouverture semilunaire, faiblement canaliculée; labre épais, lacinié, peu incurve; columelle lisse, excavée; bord columellaire étroit, calleux.” [Size small; form stocky, spire cancellate, with straight axial ribs; last whorl large, rounded at the base. Aperture semilunar, weakly canaliculate; outer lip thick, laciniate, not very curved; columella smooth, excavated; columellar margin narrow, callused.] Cossmann (1906: 134).

Discussion. The genus is characterized by its weak ventrolateral varix and the oblique aperture, convex and wide basal margin of the slightly thickened outer lip and the strongly deflected, nearly horizontal, short, twisted siphonal canal ( Figs 21A 1 –A 2). Lampanella minima has a direct development ( Kowalke 1998), which is inferred from its protoconch of 1.5 smooth, convex whorls. Its first two teleoconch whorls are moderately convex with two spiral cords (mid-whorl and close above abapical suture); third spiral cord intercalated close below adapical suture on third teleoconch whorl. Spiral cords becoming indistinctly beaded on subsequent whorls coinciding with intercalation of secondary spiral cords. The type species is very variable concerning shape, ranging from elongate shells to stockier, pupoid specimens. Similarly, its sculpture ranges from spiral cords with distinct beads to more or less prominent axial ribs but is always characterized by narrow, prominent secondary spiral threads between the primary spiral cords. As pointed out by Bequaert (1942: 8) the various morphs may co-occur and have no ecological significance (for a detailed conchological description of Lampanella minima see Bequaert 1942: 8; the description of protoconch and early teleoconch is based on Kowalke 1998 and a specimen from Florida, illustrated on [https:// www.jaxshells.org/617hh.htm; by Harry G. Lee, Dept. of Geological Sciences, University of Florida, Gainesville, USA).

Conchologically, Lampanella is surprisingly similar to the cerithiid Thericium Monterosato, 1890 and already Ozawa et al. (2019: 519) concluded: “ The small, granulose shell of Lampanella , with short anterior canal, ventrolateral varix and thickened lip is unique among batillariids, but convergent with members of the Cerithiidae . The lack of varices on the spire, lack of teeth or striae within the shell under the varices, and strongly twisted canal distinguish Lampanella from most cerithiids.” This is true indeed for “ most ” cerithiids but the absence of teeth was described by Landau et al. (2013) also as important feature to distinguish Thericium from Cerithium . Molecular data of Ozawa et al. (2019) reveal a close relationship of Lampanella with Planaxidae , which thus excludes a cerithiine affiliation and points to a striking case of convergence between Lampanella and Thericium. We note however, that Ozawa et al. (2019) did not include Thericium species in their analysis.

Stratigraphy and paleogeography. The extant Lampanella minima is distributed from Florida through the West Indies to Brazil ( Bequaert 1942). The genus appeared during the Eocene in the Western Atlantic, documented by Lampanella advena ( Palmer, 1953) from Florida. The Early Miocene Lampanella transecta ( Dall, 1890) is another record from Florida ( Dall 1890; Palmer 1953; Ozawa et al. 2009). The European record starts with Lampanella pupaeformis (de Basterot, 1825) from the Chattian of the Proto-Mediterranean Sea ( Esu & Girotti 2010) and the Early Miocene of the northeastern Atlantic ( Lozouet et al. 2001a). That species is also recorded from the Serravallian of Turkey ( Landau et al. 2013). Polymorphism and the low number of available specimens make it difficult to decide if all these occurrences represent a single species or an unresolved species complex (see Landau et al. 2013: 44). Lampanella obliquistoma ( Seguenza, 1880) represents a second European species, ranging from the Middle to Late Miocene of the Proto-Mediterranean Sea and the Central Paratethys. Despite the intraspecific variability, Lampanella pupaeformis and L. obliquistoma can be distinguished by the much higher spire and slenderer outline of L. pupaeformis . During the Sarmatian, the genus was represented by Lampanella volhynica ( Friedberg, 1914) with a restricted occurrence in the Fore-Carpathian Basin. The last Paratethyan representative of this genus is Lampanella maeotica ( Karlov, 1932) , which is restricted to the early Maeotian of the Eastern Paratethys.

Ecology. Lampanella minima lives in large populations on mud in the intertidal zone tolerating brackish waters ( Bequaert 1942). The species occurs in mangroves, but may also settle sand and rock platforms in sheltered environments ( Plaziat 1984; Ozawa et al. 2009). The tolerance for brackish water conditions may explain its occurrence in the early Maeotian of the Eastern Paratethys Sea.

Species-level taxa placed in Lampanella reported from the Paratethys attritum . Cerithium . Boettger, 1907 → Lampanella obliquistoma ( Seguenza, 1880) conoideum . Cerithium (Pithocerithium) . Karlov, 1932 → Lampanella maeotica ( Karlov, 1932) elongata. Cerithium (Pithocerithium) . Karlov, 1932 → Lampanella maeotica ( Karlov, 1932) formosum . Cerithium (Pithocerithium) . Karlov, 1932 → Lampanella maeotica ( Karlov, 1932) kerchense. Cer. (Thericium). Badzoshvili 1967 → Lampanella maeotica ( Karlov, 1932) lineata. Cerithium (Pithocerithium) . Karlov, 1932 → Lampanella maeotica ( Karlov, 1932) lineato-punctata. Cerithium (Pithocerithium) . Karlov, 1932 → Lampanella maeotica ( Karlov, 1932) maeoticum . Cerithium (Thericum) . Badzoshvili, 1967 → Lampanella maeotica ( Karlov, 1932) maeoticum . Cerithium (Pithocerithium) . Karlov, 1932 → Lampanella maeotica ( Karlov, 1932) maeoticum . Cerithium . Ossaulenko, 1936 → Lampanella maeotica ( Karlov, 1932) melaniaeformis . Cerithium (Pithocerithium) . Karlov, 1932 → Lampanella maeotica ( Karlov, 1932) obliquistoma . Cerithium . ( Seguenza, 1880) → Lampanella obliquistoma ( Seguenza, 1880) pseudobliquistoma . Pithocerithium . Szalai, 1926 → Lampanella obliquistoma ( Seguenza, 1880) punctata. Cerithium (Pithocerithium) . Karlov, 1932 → Lampanella maeotica ( Karlov, 1932) striatulum . Cerithium (Pithocerithium) . Karlov, 1932 → Lampanella maeotica ( Karlov, 1932) subpupaeformis . Cerithium . Kókay in Katona, 2011 → Lampanella obliquistoma ( Seguenza, 1880) volhynicum . Cerithium . Friedberg 1914 → Lampanella volhynica ( Friedberg, 1914)