Geostiba (Sibiota) aucta, Assing, 2006

Assing, V., 2006, Thirteen new species and additional records of Eastern Mediterranean Geostiba THOMSON (Coleoptera: Staphylinidae, Aleocharinae), Linzer biologische Beiträge 38 (2), pp. 1179-1215 : 1179-1215

Geostiba (Sibiota) aucta sp.n. ( Figs 86-97 , Map 8 )

Holotype 3: TR [24] - Rize, 25 km SSE Rize, 7 km E Ikizdere , 1030 m, 40°47'01N, 40°38'18E, 31.VII.2006, V. Assing / Holotypus 3 Geostiba aucta sp.n. det. V. Assing 2006 (cAss). Paratypes: 233, 1♀: same data as holotype (cAss) ; 8 exs.: same data, but leg. Schülke (cSch, OÖLL) ; 2♀♀: TR [23] - Rize, 25 km SSE Rize, 4 km E Ikizdere , 750 m, 40°47'14N, 40°35'31E, 31. VII.2006, V. Assing (cAss) ; 2 exs., same data, but leg. Schülke (cSch) ; 13: same data, but "[23a] ... 1.VIII.2006, V. Assing" (cAss) ; 3 exs.: same data, but leg. Schülke (cSch) .

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D e s c r i p t i o n: Moderately small species, 2.4-3.0 mm (abdomen fully extended). Coloration: body reddish brown to brown, with abdominal segment VI and adjacent segments more or less distinctly infuscate and sometimes also the head slightly darker; legs yellowish; antennae yellowish brown.

Facies as in Fig. 86 . Head with very shallow microreticulation and some shine ( Fig. 87 ); puncturation very fine and very sparse, barely noticeable. Eyes small ( Fig. 88 ), not projecting from lateral outline of head, approximately 1/4 the length of postocular region in dorsal view.

Pronotum weakly transverse, approximately 1.1 times as wide as long and 1.2 times as wide as head ( Fig. 87 ); microsculpture similar to that of head or slightly more pronounced; puncturation similar to that of head.

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Elytra approximately 0.6 times as long as pronotum ( Fig. 87 ), with pronounced sexual dimorphism; microsculpture similar to that of head and pronotum; puncturation weakly granulose.

Abdomen approximately 1.2 times as wide as elytra ( Fig. 86 ), with distinct microsculpture, and with sparse fine puncturation; posterior margin of tergite VII without palisade fringe.

3: elytral disc with pronounced impression, on either side of suture distinctly elevated, these elevation broader anteriorly than posteriorly and granulosely sculptured ( Fig. 89 ); tergite VII near posterior margin with pair of short and rather weakly elevated carinae ( Fig. 90 ); posterior margin of tergite VIII in the middle almost truncate and with only few setae ( Fig. 91 ); posterior margin of sternite VIII weakly convex; median lobe of aedeagus with reduced crista apicalis and with series of semitransparent spines in internal sac ( Fig. 92 ); apical lobe of paramere as in Fig. 93 .

♀: elytra very shallowly impressed; posterior margin of tergite VIII weakly convex ( Fig. 94 ); posterior margin of sternite VIII weakly convex and with row of numerous modified marginal seta, in the middle not distinctly concave ( Fig. 96 ); spermatheca with distinctly dilated capsule ( Figs 96-97 ).

E t y m o l o g y: The name (Lat., adj.: enlarged) alludes to the shape of the capsule of the spermatheca, one of the characters distinguishing this species from G. rizensis .

C o m p a r a t i v e n o t e s: In general appearance, the new species is highly similar to the parapatric G. rizensis , its geographically closest consubgener. From this species, G. aucta is separated especially by the broader, more granulosely sculptured, and less sharply delimited sutural elevations on the male elytra, the (on average) less pronounced carinae on the male tergite VII, by the reduced crista apicalis of the median lobe (in G. rizensis pronounced), the more numerous and shorter spines in the internal sac, the different shape and chaetotaxy of the apical lobe of the paramere (in G. rizensis with one very long median and three much shorter apical and subapical setae), as well as by the much larger spermathecal capsule. For illustrations of the aedeagus and the spermatheca of G. rizensis see PACE (1983) and ASSING (2001a). In order to account for G. aucta , the key in ASSING (2005a) is modified as follows: 100. 3: sutural carinae anteriorly narrower, less strongly elevated, and without coarse puncturation or sculpture; tergite VII with usually pronounced, though relatively short pair of carinae, these carinae separated by a distance distinctly less than their width and posteriorly merging (Fig. A01a: 56); median lobe of aedeagus apically more acute (ventral view) and less slender (Figs. A01a: 51-52). ♀: spermatheca as in Figs. A01a: 54-55. Northeastern Anatolia: Artvin ................. G. fabaeformis ASSING - 3: sutural carinae anteriorly wider and strongly elevated, decreasing in width and elevation posteriad, coarsely and granulosely sculptured, and mat; pair of carinae at posterior margin of tergite VII almost straight and parallel, sometimes separated by a distance greater than their width ( Figs 90 , A01a: 59). Northeastern Anatolia:

Rize .........................................................................................................................100a 100a. 3: sutural carinae narrower, less granulosely sculptured, and more sharply delimited; median lobe of aedeagus with pronounced crista apicalis and with fewer and longer spines in internal sac ( Figs P83b : 90-91); apical lobe of paramere with one very long basal and three much shorter subapical and apical setae. ♀:

spermatheca with slender capsule of conical shape (Fig. A01a: 58). Rize: area to the south of Ikizdere............................................................................. G. rizensis PACE

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- 3: sutural carinae broader, more granulosely sculptured, and less sharply delimited ( Figs 87, 89 ); median lobe of aedeagus with reduced crista apicalis and with more numerous and shorter spines in internal sac ( Fig. 92 ); apical lobe of paramere as in Fig. 93 , with long subapical and apical setae. ♀: spermathecal capsule distinctly enlarged ( Figs 96-97 ). Rize: area to the east of Ikizdere ( Map 8 )............ G. aucta sp.n.

D i s t r i b u t i o n a n d b i o n o m i c s: Thetypespecimenswerefound in two localities in the area to the east of Ikizdere ( Map 8 ). They were collected by sifting the leaf litter in mixed forests with chestnut, alder, beech and Rhododendron at altitudes of 750 and 1030 m.

Figures

Map 8: Distributions of Geostiba aucta (filled circles), G. orduica (square), and G. soganlica (open circle) in Turkey.

Figs 86-97: Geostiba aucta sp.n.: (86) 3 habitus; (87) 3 forebody; (88) head in lateral view; (89) 3 elytra in dorso-lateral view; (90) 3 abdominal tergite VII in dorsal view; (91) 3 tergite VIII; (92) median lobe of aedeagus in lateral view; (93) apical lobe of paramere; (94) posterior margin of ♀ tergite VIII; (95) posterior margin of ♀ sternite VIII; (96-97) spermatheca. Scale bars: 86: 1.0 mm; 87: 0.5 mm; 88-91, 94-95: 0.2 mm; 92-93, 96-97: 0.1 mm.

Figs 78-84: Geostiba priva sp.n.: (78) 3 habitus; (79) 3 forebody; (80) head in lateral view; (81) 3 abdominal tergites VII-VIII in lateral view; (82) 3 tergite VII in antero-dorsal view; (83) median lobe of aedeagus in lateral view; (84) spermatheca. Scale bars: 78: 1.0 mm; 79-82: 0.2 mm; 83-84: 0.1 mm.