Clistoabdominalis namibiensis, FÖLDVÁRI, MIHÁLY, 2013

FÖLDVÁRI, MIHÁLY, 2013, Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae), Zootaxa 3656 (1), pp. 1-121 : 21

Clistoabdominalis namibiensis sp. nov.

( Figs 6G–L , 39N–O )

Diagnosis: The species is unique, because of the flat, spiny lobe of the OG and the visible S7. Other special features are the two subapical (distal) spines on mid tibiae of both sexes and the cross-vein R-M being placed at 1/ 5 of the discal cell.

Type material: Namibia: 1♂, HT [# T645], Omaruru dist[rict], Ugab R [iver] n[ea]r Brandberg Wes [t], 20°57’39’’S 14°07’55’’E, 21–24.x.1998, Kirk-Spriggs & Marais, Malaise trap sample ; 1♀, AT, same data as HT (both NMNW) ; 1♂, 1♀, PTs, same data as HT (both HNHM) ; 1♀, PT, Khorixas district , Huab River, Krone 721, 20°37’09’’S 13°54’31’’E, 23–26.x.1998, Kirk-Spriggs & Marais, Malaise trap ( NMNW) .

Other material examined: South Africa: 1♂, Cape Prov, Papendorp, at mouth of Olifants River , Sept. 11 1972, sea level, 3118 Cc, ME. Irwin, Coastal dunes; Cape Prov, 12 mi NNE. Garies, 3018 Ac, Sept 9 1972, M.E. Irwin, 1350ft, nr. waterfall (both NMSA) .

Male

Head. Third antennal segment acuminate; yellow, whitish towards tip. Face silvery pollinose. Frons, upper part shining black, lower part silvery grey; eyes touching for distance equal to 2–2.5 times ocellar triangle. Occiput silvery pollinose.

Thorax. Humeri pale yellow. Mesonotum (viewed obliquely from front) silvery pollinose, also from the side. Scutellum grey, hairs missing. Dorsocentral hairs distinct. Halter whitish yellow, brown at base of stem.

Legs. Trochanters and base of femora yellow brown, femora yellow with darker ring in the middle (3/5), knees and tibiae yellow. Tarsal segments yellow, last segment brown. Ventroapical row of 6–7 tiny, dark spines on mid femora only. Subapical (distal) spines on first four tibiae present (two spines on t2). One very weak anteromedial hair on 3rd tibia. Hind trochanter with a few short, pale hairs on ventral side. Pulvilli as long as last tarsal segment. Wing. Fourth costal section as long as third costal section. Cross-vein R-M at 1/5 of discal cell. Pterostigma fully coloured. Hairs on tegula missing.

Abdomen. Viewed obliquely from front tergites brownish pollinose, with grey patches laterally in dorsal view, hind margins brown (distal 1/3 of T5 silvery pollinose), sides greyish. Hairs dispersed, short and weakly developed. Laterally 2–3 short spines on first tergite. Postabdomen in dorsal view: S7 visible; T5 as long as ST8. Genitalia without dissection: the whole postabdomen yellow-brown, ST8 swollen, m.a. slit-like, oblique.

Genitalia. Surstyli asymmetrical, is larger; epandrium broad, u-shaped around cerci; ST8 enlarged ( Fig. 6K ). SES clearly developed, without distinct hairs; gonopods highly asymmetrical, OG bearing a broad, flat, spiny lobe, ig hump-like; lobes of hypandrium rectangular ( Fig. 6J ); phallic guide with parallel sides, narrowing at tip; phallus trifid, with supporting membranous structures, branches pointed at tips; ejaculatory apodeme triangle shaped; sperm pump flat, with two projections ( Fig. 6I ).

Female

As male except for the following characters. Frons, eyes separated; silver-grey pollinose on lower half otherwise shining black. Pulvilli and claws about 1.5–2 times as long as last tarsal segment. Female ovipositor ( Figs 39N–o ): base enlarged, hemispherical, yellow-brown; S6 modified to receive piercer.

Distribution— Namibia, South Africa.

Etymology —The species is named after the African country Namibia.

Figures

FIGURE 6. Male genitalia of Clistoabdominalis lomholdti (A–F) and Cli. namibiensis sp. nov. (G–L).—A: outer surstylus with phallic guide and outer gonopod, lateral view, B: dorsal view, C: ventral view with phallic guide and phallus (without ST8), D: surstyli, dorsal view, E: inner surstylus, lateral view, F: sperm pump with ejaculatory apodeme, G: outer surstylus with phallic guide, phallus and outer gonopod, lateral view, H: inner surstylus, lateral view, I: sperm pump with ejaculatory apodeme, J: ventral view with phallic guide and gonopods (without ST8), K: dorsal view, L: surstyli, dorsal view. Scale bars: 0.1 mm

FIGURE 39. Female ovipositors.—A: Eudorylas liberia, B: Eudorylas setiformis, C: Eudorylas aemulus, D: Eudorylas garambensis, E: Eudorylas remiformis, F: Eudorylas mutillatus (E. eremnoptera), G–H: Eudorylas definitus (E. excisus), I: Eudorylas facetus, J–K: Dasydorylas quasidorsalis, L: Eudorylas amuscarinum, M: Eudorylas abdominalis, N–O: Clistoabdominalis namibiensis sp. nov., P: Eudorylas mutillatus, Q: Eudorylas femoralis sp. nov., R: Eudorylas ghesquierei, S: Dasydorylas okongoensis sp. nov., T–U: Eudorylas angustus, V: Eudorylas excisus. Scale bar: 0.5 mm