Helix (Helix) philibinensis ROSSMÄSSLER, 1839

Neubert, Eike, 2014, Revision of Helix LINNAEUS, 1758 in its eastern Mediterranean distribution area, and reassignment of Helix godetiana KOBELT, 1878 to Maltzanella HESSE, 1917 (Gastropoda, Pulmonata, Helicidae), Contributions to Natural History 26, pp. 1-200 : 57-64

Helix (Helix) philibinensis ROSSMÄSSLER, 1839 ( Figs 89–96 )

1839 Helix philibinensis ROSSMÄSSLER, Iconographie der Land- und Süsswassermollusken 1 (2) (3/4): 10, Taf. 44, Fig. 581 [Macedonien].

1901 Pomatia ( ligata subsp.) oestreichi KOBELT, Iconographie der Land- und Süsswassermollusken 2 (9): 54, Taf. 258, Fig.1674 [in der Gegend des Presba Sees in Albania].

1904 Helix frivaldszkyi KOBELT, Systematisches Conchylien-Cabinet VI : 146, Taf. 338, Fig. 8, 9 [ Türkei, wahrscheinlich in Makedonien oder im benachbarten Teil von Rumelien].

1906 Helix (Helicogena) despotina KOBELT, Systematisches Conchylien-Cabinet VI : 279, Taf. 371, Fig. 9 , 10 [Rhodope Gebirge (Despoto Dagh) gegen Philippopel (= Plovdiv)].

1906 Helix (Helicogena) thracica KOBELT, Systematisches Conchylien-Cabinet VI : 269, Taf. 368, Fig. 5, 6 [Rhodope Gebirge (Despoto Dagh) gegen Philippopel].

1906 Helix (Helicogena) albescens bulgarica KOBELT, Systematisches Conchylien-Cabinet VI : 275, Taf. 370, Fig. 7–10 [Rhodope Gebirge (Despoto Dagh) gegen Philippopel].

1907 Helix (Helicogena) speideli O. BOETTGER, Nachrichtsblatt der deutschen Malakozoologischen Gesellschaft 39: 37 [Thasos].

Type specimens:

philibinensis ( = frivaldszkyi fide Zilch 1952): holotype frivaldszkyi SMF 9775 ( Zilch 1952: 160) H = 28.0; D = 27.4; PH = 19.7; PD = 14.4; PrD = 6.2; W = 4.5. oestreichi: holotype SMF 9616 ( Zilch 1952: 159): H = 22.5; D = 28.6; PH = 19.5; PD = 14.9; PrD = 5; W = 4.5. despotina : holotype SMF 9659 ( Zilch 1952: 161): H = 28.1; D = 25.4; PH = 20.3; PD = 12.6; PrD = 5.7; W = 4.5. thracica : H =; D =; PH =; PD =; PrD =; W =. bulgarica: lectotype SMF 9685 (SD Zilch 1952: 161): H = 29.1; D = 30.2; PH = 21; PD = 15; PrD = 6; W = 4.5. speideli : lectotype SMF 21545 (SD Zilch 1952: 161): H = 33.8; D = 31; PH = 23.1; PD = 15.9; PrD = 6.1; W = 4.75.

Specimens examined:

Bulgaria: SMF 74390/2 Sofia, 42.6907 23.3227, coll. Jetschin ex Nägele; SMF 158641, Plovdiv, hill of Bunardzik, 42.1444 24.7384, coll. Zilch ex Urbanski; SMF 158904/3, Assenova Krepot close to Assenovgrad, 41.9876 24.8756, coll. Zilch ex Urbanski; SMF 158642/2 Assenovgrad in the Rhodopes, coll. Zilch ex Urbanski; SMF 86141/6, Plovdiv, Sacháte Tepe, coll. Ehrmann ex Matschewa; Pirin Mts, on the western slope of Sveštenik Mt., Kalimantsi, Peshternik area , limestone gorge, 41.5166 23.6333, 08.06.2006, leg. I. Dedov, NMBE 519950 /2 (alk.). Not checked: Plindenci, ca. 20 km N Sandanski (= 41.5502 23.2456) (( Hudec & Vasatko (1973)).

FYROM: SMF 86142/9 NE of the Dojran Lake, 41.2554 22.7114, ex RÖmheld 1916; ditto, SMF 86143 / 2; SMF 22654/1 at the Vardar, ex RÖmheld 1916; Demir Kapija at the Vardar, 108 m alt., 41.41 22.28, 29.7.1984, NMBE 512565 / 2; ditto, 3.7.1986, NMBE 512566 / 2; road Carina-Sveti Naum, Galicica Mts. , 40.9718 20.8788, 1000 m alt., 14.7.1972, NMBE 524472 / 1; Belovodica in dir. Gradsko, hill slope, 41.3508 21.708, 710 m alt., 11.7.1972, NMBE 524471 / 3; crossroads from Demir Kapija, 3.5 Km S in dir. Gevgelija, 41.4087 22. 2826, 105 m alt., 28.7.1990, NMBE 524464758 .

Greece: Thessaloniki, Langadas , 40.7513 23.0683, coll. S. H. Jaeckel ex Haltenroth, SMF 214892 ; Serres, Akropolis , 41.0909 23.5413, 22.09.1984, LIEB (13) ; Xanthi , 41.1367 24.8906, 22.06.1989, LIEB (4) ; Makedonia, ruins of Philippi , 50 m alt., 41.0093 24.2798, 30.7.1984, NMBE 512564 / 1; N of the Vegoritsas Lake 1 km W of Arnissa, 800 m alt., 40.804 21.755, 1.6.1987, NMBE 5125672 / 1; Mikra Prespa at Mikro Limni , 860 m alt., 40.743 21.11, 17.7.1990, NMBE 512568 / 4; isthmus at Mikra Prespa at Mikro Limni , 860 m alt., 40.805 21.0714, 1986, NMBE 512572 / 10; Kastoria , rocks on the peninsula, 640 m alt., 40.505 21.279, 17.7.1990, NMBE 512569 / 4; Kavala, summit area of the Pangäon , 40.914 24.09, 1930 m alt., 8.9.1991, NMBE 512571 / 1; Stavroupoli, close to Nestos shore, 41.2031 24. 7056, 120 m alt., 12.5.2008, NMBE 519953 / 1 (alk.); Zarkadia , 2.5km along the road, 41.0177 24. 6405, 140 m alt., 22.9.1989, NMBE 519952 / 2 (alk.).; Evvia, Mantoudi , 38.7942 23.477, 20 m alt., 21.5.1997, NMBE 524590 / 7; Amarantos, Kalkberg E von Loutra, 40.1745 20.7297, 1250 m alt., 16.5.1997, NMBE 524609 / 1; Eleohoria, 500 m N/ Chalkidiki Peninsula , 40.3434 23. 1613, 170 m alt., 5.5.1995, NMBE 524623 / 1; Petralona , cave, 40.3721 23. 1685, 335 m alt., 29.5.1997, NMBE 524624 / 3; ditto, 5.5.1995, NMBE 524425 / 2; Gérmas , valley W of village, 40.4528 21. 4149, 850 m alt., 17.5.1997, NMBE 524595 / 4; road Saloniki-Galatista , 9 km E Galatista, 40.465 23.364, 460 m alt., 28.8.1991, NMBE 512570 / 1; Monastir Panagia Mavriotissa , 40.505 21. 2786, 640 m alt., 14.7.1990, NMBE 524463 / 10; Nymfopetra, W Volvi Lake , 40.6922 23.3085, 70 m alt., 24.9.1989, NMBE 524466 / 5; 4.5 Km W crossroads to Loutra Elefthero, 40.7211 24.0601, 30 m alt., 21.9.1989, NMBE 524462 / 50; Mikra Prespa Lake , 200 m W Mikrolimni, 40.7419 21. 1078, 885 m alt., 13.7.1990, NMBE 524451 / 5; Mikrolimni , W, 40.7426 21. 1098, 870 m alt., 10.5.1995, NMBE 524433 / 1; pass close Edessa, 1 km to Arnissa, Vegoritses Lake , 40.804 21.755, 800 m alt., 21.6.1987, NMBE 512567 / 1; Galipsos , 40.8062 23.9502, 200 m alt., 6.5.1995, NMBE 524434 / 2; Megali- and Mikra Prespa Lakes , confluent, 40.8111 21.0706, 855 m alt., 13.7.1990, NMBE 524456 / 19; Psarades , 40.8315 21.0288, 857 m alt., 10.5.1995, NMBE 524435 / 2; Sidirohori, 1.2 Km in dir. Folia , Simvolo Mts. , 40.838 24. 1375, 250 m alt., 27.5.1997, NMBE 524588 / 7; mountains N Podohori , 40.8473 24.0249, 450 m alt., 27.5.1997, NMBE 524602 / 1; Platanotopos, 1 Km in dir. Eleftheroupoli , 40.8524 24.0717, 280 m alt., 27.5.1997, NMBE 524578 / 3; Mousthéni , 40.8566 24. 1137, 175 m alt., 6.5.1995, NMBE 524444 / 11; road Aridéa-Skidra, crossroads to Néa Zoi , 40.8629 22.1183, 200 m alt., 15.5.2008, NMBE 523313 / 17; Akrovouni, 16.5 Km, Pangéo Mts. , 40.9094 24.1406, 1350 m alt., 6.5.1995, NMBE 524587 / 1; Pangéo summit, 9.2 Km in dir. Akrovouni , "Katafigi", 40.9109 24.1414, 1350 m alt., 27.5.1997, NMBE 524619 / 5, NMBE 524606 / 1; Akrovouni, 25.2 Km in dir. Pangéo summit, 40.9113 24.0973, 1780 m alt., 23.9.1989, NMBE 524457 / 9; Akrovouni, 17.8 Km in dir. Pangéo summit, 40.9147 24.1465, 1280 m alt., 23.9.1989, NMBE 524454 / 7; Akrovouni, 11.6 Km in dir. Pangéo summit, 40.9171 24.1906, 900 m alt., 23.9.1989, NMBE 524469 / 1; Akrovouni, little canyon 8 Km N Akrovouni , 40.921 24.206, 600 m alt., 6.5.1995, NMBE 524586 / 2; Pangéo summit, 1.3 Km in dir. Akrovouni , 40.9122 24.0936, 1850 m alt., 23.9.1989, NMBE 524460 / 3; Pangéo summit, below radio station, 40.9138 24.0903, 1950 m alt., 23.9.1989, NMBE 524459 / 1; Akrovouni, 12.3 Km in dir. Pangéo summit, 40.9222 24. 1876, 950 m alt., 23.9.1989, NMBE 524470 / 1; Akrovouni, 7.5 Km in dir. Pangéo summit, 40.9237 24. 2178, 480 m alt., 27.5.1997, NMBE 524620 / 3; Nikissiani , 40.9462 24.144, 355 m alt., 6.5.1997, NMBE 524617 / 5; Nikissiani , S of village, 40.9467 24. 1439, 350 m alt., 22.9.1989, NMBE 524618 / 2, NMBE 524461 / 2; Kavala, 2.5 Km E, 40.9486 24.4438, 20 m alt., 22.9.1989, NMBE 524465 / 17; Griva, Gouménissa , 40.9562 22. 4009, 540 m alt., 20.5.1997, NMBE 524580 / 3; ditto, 13.5.2005, NMBE 523314 / 18; ditto, 14.5.2008, NMBE 523317 / 1; Griva , canyon, 40.9574 22. 4055, 480 m alt., 9.5.1995, NMBE 524585 / 2; Griva, 4.5 Km NW canyon, 40.9709 22.3769, 600 m alt., 19.4.2009, NMBE 523316 / 7; below Moni Ikosifinissi , 40.9652 24. 1064, 575 m alt., 6.5.1995, NMBE 524455 / 1; Loutra Arides, Taleingang , 40.9722 21.912, 450 m alt., 19.5.1997, NMBE 524429 / 29; ditto, 14.5.2008, NMBE 523315 / 7; ditto, 26.5.2011, NMBE 523299 / 6; Loutraki , valley behind village, 40.9769 21. 9454, 350 m alt., 19.5.1997, NMBE 524603 / 12; Palea Kavala, 4.2 Km in dir. Halkero , 40.9994 24. 4273, 440 m alt., 26.5.1997, NMBE 524583 / 3; Palea Kavala, 3 Km W of village, 41.0009 24.3934, 200 m alt., 7.5.1995, NMBE 524436 / 1; Philippi, Krinides , above the ruin, 41.0128 24.2851, 85 m alt., 7.5.1995, NMBE 524467 / 2; Palea Kavala, 2.5 Km in dir. Korifés , 41.017 24.4388, 500 m alt., 7.5.1995, NMBE 524442 / 4; Alistrati, canyon 6.5 Km SE village, 41.0224 24.0067, 80 m alt., 23.5.1997, NMBE 524604 / 20; Zarkadia, 2.5 Km NNE, 41.0288 24. 6417, 450 m alt., 22.9.1989, NMBE 524450 / 6, NMBE 524424 / 1; Mt. Zarkadia , 41.0293 24. 6415, 470 m alt., 7.5.1995, NMBE 524439 / 3; Makrihori, valley 2.5 Km E in dir. Kehrokambos , 41.0466 24. 6462, 180 m alt., 26.5.1997, NMBE 524615 / 5; Kehrokambos, 7.2 Km in dir. Makrihori , 41.1043 24. 6392, 550 m alt., 26.5.1997, NMBE 524605 / 3; Kehrokambos , 400 m S village, 41.1514 24. 6454, 390 m alt., 7.5.1995, NMBE 524430 / 7; Kehrokambos, 4 Km in dir. Stavroupoli , 41.1806 24. 6687, 180 m alt., 7.5.1995, NMBE 524468 / 1; Neohori , canyon entrance, 41.0481 22.0893, 275 m alt., 9.5.1995, NMBE 524438 / 13; ditto, 19.5.1997, NMBE 524589 / 21; ditto, 27.5.2011, NMBE 523295 / 4; Limnia, 5 Km in dir. Zigos , 41.0589 24. 3816, 730 m alt., 26.5.1997, NMBE 524613 / 3; mountain E Makrihori , 41.0692 24. 6298, 650 m alt., 7.5.1995, NMBE 524431 / 1; Agios Kosmas 1 Km to Makrihori , 41.0783 24. 6536, 380 m alt., 26.5.1997, NMBE 524611 / 3; Paradissos , NE village, 41.0843 24.7612, 60 m alt., 25.5.1997, NMBE 524592 / 3; Platamonas , 41.1001 24. 5301, 720 m alt., 26.5.1997, NMBE 524610 / 2; Inoussa , 41.1059 23.6312, 200 m alt., 21.5.1997, NMBE 524593 / 5; Notia , 41.1072 22. 2115, 650 m alt., 9.5.1995, NMBE 524427 / 7; Likostomo , 41.1078 24. 4424, 740 m alt., 26.5.1997, NMBE 524614 / 4; Agora, 1.3 Km in dir. Peristéria , 41.1165 24. 3244, 380 m alt., 23.5.1997, NMBE 524581 / 9; Peristéria, 1 Km W in dir. Adriani , 41.1177 24.3837, 400 m alt., 23.5.1997, NMBE 524607 / 1; Dipotamos , 800 m W village, 41.1196 24. 5762, 690 m alt., 26.5.1997, NMBE 524445 / 5; Amaxades, 6.3 Km in dir. Iasmos , 41.1204 25.1406, 60 m alt., 27.7.1990, NMBE 524453 / 9; canyon 500 m from Kalithia to Anthohóri , 41.1249 23. 9293, 215 m alt., 8.5.1995, NMBE 524446 / 5; Xanthi , 2 km N city, 41.1591 24. 8806, 135 m alt., 27.7.1990, NMBE 524452 / 2; Nikiforos, bridge 1.8 Km E village, 41.1726 24. 3345, 235 m alt., 23.5.1997, NMBE 524591 / 6; Xiropotamos , 41.1951 24. 1002, 285 m alt., 8.5.1995, NMBE 524437 / 4; Petroussa , canyon NW village, 41.2014 24.015, 285 m alt., 13.5.2008, NMBE 523318 / 1; Stavroupoli, shore of Nestos River , 41.2031 24. 7056, 120 m alt., 27.7.1990, NMBE 524458 / 16; ditto, 24.5.1997, NMBE 524440 / 7; ditto, 12 5.2008, NMBE 523300 / 10; Makriplagio, 2.6 Km SW, 41.2082 24. 2519, 520 m alt., 8.5.1995, NMBE 524441 / 3; ditto, 22.5.1997, NMBE 524601 / 2; road Xanthi-Stavroupoli, 2 Km in dir. Ehinos , 41.2085 24.8612, 200 m alt., 7.5.1995, NMBE 524426 / 11; ditto, 24.5.1997, NMBE 524608 / 1; ditto, 12.5.2008, NMBE 523294 / 4; Prossotsani, 6 Km N village, 41.2225 23. 9694, 340 m alt., 8.5.1995, NMBE 524447 / 10; ditto, 13.5.2008, NMBE 523297 / 4; 25.5.2011, NMBE 523296 / 3; Prossotsani, 8 Km in dir. Granitis , 41.2597 23.9737, 500 m alt., 22.5.1997, NMBE 524582 / 4; Makriplagio , 41.2267 24. 2713, 880 m alt., 22.5.1997, NMBE 524612 / 3; Taxiarhis , 41.2328 24.1888, 500 m alt., 22.5.1997, NMBE 524579 / 7; ditto, 12.5.2008, NMBE 523298 / 7; Petroussa, canyon 3–3.5 Km in dir. Pirgi , 41.2332 24.0135, 510 m alt., 21.5.1997, NMBE 524596 / 2; Makriplagio, 3.4 Km NE village, 41.2333 24.3013, 800 m alt., 8.5.1995, NMBE 524432 / 1; Sidirokastro , above city, 41.2398 23. 3954, 110 m alt., 20.5.1997, NMBE 524584 / 28; Panorama , above city, 41.2511 23. 8208, 650 m alt., 21.5.1997, NMBE 524594 / 11; Kato Vrontou, 4.3 Km in dir. Panorama , 41.262 23. 7986, 750 m alt., 21.5.1997, NMBE 524622 / 1; road Prossotsani-Granitis , 41.2821 23. 9815, 680 m alt., 8.5.1995, NMBE 524443 / 2; Kato Nevrokopi , 41.3498 23.881, 650 m alt., 21.5.1997, NMBE 524600 / 2; Sidironero , hill slope NE village, 41.3686 24. 2339, 650 m alt., 22.5.1997, NMBE 524621 / 1; Mikroklissoura, canyon 1 Km in dir. Potami , 41.3867 24.0698, 400 m alt., 22.5.1997, NMBE 524597 / 1; Ahladia, 2.9 Km in dir. Kato Nevrokopi , 41.3885 23. 9748, 690 m alt., 22.5.1997, NMBE 524428 / 1. Thasos: Island of Thasos , coll. O. Boettger ex Speidel (paralectotypes of speideli ), SMF 9839 / 2; ditto, SMF 9941 / 7 (paralectotypes of speideli ); ditto, SMF 74388 / 3, coll. Jetschin ex C.R. Boettger (paralectotypes of speideli ?); ditto, SMF 74389 / 5, coll. C.R. Boettger (paralectotypes of speideli ?); ditto, coll. Nägele ex O. Boettger, SMF 76883 / 5 (paralectotypes of speideli ); Island of Thasos 1988, coll. Hohorst ex H.G. Beckmann, SMF 311936 / 12; Island of Thasos, Theologos to Ypsarion , 40.6767 24.6791, 29. 09.–13.10.1988, coll. Hohorst ex H.G. Beckmann, SMF 311937 / 6, SMF 311938 / 1 + SMF 311939/3; Island of Thasos, slope of the Ypsarion Mountains , 40.7254 24.6754, 26.09.1991, SMF 311940 / 2; Theologos , 40.6591 24.6941, 04.09.1991, coll. Hohorst ex H.G. Beckmann, SMF 311941 / 1; Kalivia, 15 Km in dir. Aliki , 40.5914 24. 6683, 110 m alt., 21.9.1989, NMBE 524599 / 1, NMBE 524448 / 8; Aliki , S of village, olive plantation, 40.605 24.7416, 21.9.1989, NMBE 524417 / 12; Theologos , above cemetery, 40.6634 24.7022, 300 m alt., 21.9.1989, NMBE 524616 / 2; Theologos, 2.7 Km NE village, small canyon, 40.6685 24.732, 500 m alt., 21.9.1989, NMBE 524598 / 1; Theologos, 2.8 Km NE village, 40.6696 24. 7329, 530 m alt., 25.7.1990, NMBE 524449 /11.

Turkey: SMF 86144 Adrianopel (= Edirne), 41.6818 26.5622, coll. Kobelt ex Wohlberedt.

Diagnosis: shell of medium size, protoconch large, five distinct brown spiral bands, labial callus cream to brown, umbilicus closed; epiphallus short, long flagellum, short diverticulum.

Description: shell of medium size, thin, elongate spherical, with the spire high conical; basic shell colour greyish white with chestnut brown spirals; protoconch large in comparison to shell, its diameter ranging from 5–6 mm, smooth, white whorls; teleoconch of 3–4 whorls; usually with five distinct brown spiral bands; sometimes spiral No. 2 fused with No. 3; last whorl only slightly descending below the periphery; aperture subquadrate to rounded, apertural rim narrow, labial callus weak, cream to brown coloured, columellar triangle small; umbilicus closed in adult shells.

Genital organs ( Fig. 95 ): penis club-shaped, epiphallus considerably short- er than penis; flagellum extremely long, exceeding the length of epiphallus+penis fivefold; mrp attaching in a central position on the epiphallus; internally, both papillae of equal size, penial chamber long, filled with a few ring-like pilasters crossed by few elongate folds, both papillae conical with a central perforation; atrial stimulator a rounded knob; female system with a very short vaginal tube, dart sac and glandulae mucosae strongly developed, the glandulae with a basic stem, multi-branched, longer than dart sac; pedunculus stem long, with a thick and short diverticulum, bursa copulatrix stem shorter than pedunculus stem, vesicle rounded and bent downwards.

Distribution ( Fig. 96 ): This species is only known from a small area in the Central Balkan, from the region around the Ohrid Lake to the east as far as the Island of Thasos. Interestingly, it inhabits a wide range of habitats, from the lowlands up to mountain peaks as for example the Pangäon Mt., and summits in the Pirin Mt. in Bulgaria, which reach almost 2000 m altitude.

Remarks: Helix philibinensis can be discriminated from the similarly sized H. figulina by its brownish aperture (always white in H. figulina ). It differs from H. albescens by its somewhat more elongate spire and the less ribbed shell, and anatomically by the presence of a diverticulum (missing in H. albescens ). The shells of H. pomacella are considerably smaller and more globose.

Figures

Figs 89–94. Helix (Helix) philibinensis ROSSMÄSSLER, 1839. Fig. 89 philibinensis (= frivaldszkyi), holotype frivaldszkyi SMF 9775, Macedonien, D = 28.0; Fig. 90 holotype oestreichi SMF 9616, in der Gegend des Presba Sees in Albania, D = 28.6; Fig. 91 holotype despotina SMF 9659, Rhodope Gebirge gegen Philippopel, D = 25.4; Fig. 92 bulgarica: lectotype SMF 9685, Rhodope Gebirge gegen Philippopel, D = 30.2; Fig. 93 speideli lectotype SMF 21545, Thasos, D = 31.0; Fig. 94 NMBE 519950, Sveštenik Mt., Kalimantsi, D = 30.05. — All photos Neubert, original size.

Fig. 95. Helix (Helix) philibinensis ROSSMÄSSLER, 1839. Genital organs. NMBE 519950, Sveštenik Mt., Kalimantsi, situs 55.57 — All photos Neubert.

Fig. 96. Helix (Helix) philibinensis ROSSMÄSSLER, 1839. Distribution.

Figs 1–9. Helix (Helix) pomatia LINNAEUS, 1758. Fig. 1 compacta, syntype SMF 9573, D = 50.07, Hungary "Gestrüppe der Berglehnen"; Fig. 2 pulskyana, syntype SMF 9525/2a, Hungary "Inseln der Donau", D = 36.6; Fig. 3 hajnaldiana, syntype SMF 9658, Hungary "Budapest, im botanischen Garten", D = 34.1; Fig. 4 solitaria, syntype SMF 74708/2a, D = 31.1; Fig. 5 sabulosa, syntype SMF 9524, D = 40.5, Fig. 6 expansilabris, holotype SMF 9538, D = 45.5; Fig. 7 luteola, holotype SMF 9527, D = 34.6; Fig. 8 pannonica, lectotype SMF 9535, D = 40.5, Fig. 9 lednicensis, syntype SMF 9534/2a, D = 38.3. — All photos Neubert, original size.

Figs 10–19. Helix (Helix) pomatia LINNAEUS, 1758. Fig. 10 claudiensis, holotype SMF 9543, Klausenburg, D = 39.5; Fig. 11 dobrudschae, holotype SMF 9541, Rumania, Dobrudscha, Comana Vlasca, D = 46.2; Fig. 12 costellata, holotype SMF 9631, Markt in Bukarest, D = 39.7; Fig. 13 elsae, lectotype SMF 9529, nahe Bukarest, D = 34.9; Fig. 14 transsylvanica, holotype SMF 9634, Markt in Bukarest, D = 40.1; Fig. 15 christinae, lectotype SMF 9542, Runku über Bukarest, D = 44.5; Fig. 16 kapellae, holotype SMF 9544, Kapella Gebirge, D = 44.2; Fig. 17 serbica, holotype SMF 9540, Maidanek, D = 53.9; Fig. 18 thessalica, lectotype SMF 9568, Makrinitza bei Volos, D = 45.4; Fig. 19 rhodopensis, syntype SMF 9599/2a, Philippoppel, D = 41.0. — All photos Neubert, original size.