Agra catbellae Erwin

Erwin, Terry L., 2002, The Beetle Family Carabidae of Costa Rica: Twenty­nine new species of Agra Fabricius 1801 (Coleoptera: Carabidae, Lebiini, Agrina), Zootaxa 119, pp. 1-68 : 56-58

Agra catbellae Erwin , new species

( Figs. 25a, 25 b, 25c, 25d)

Holotype. Male, COSTA RICA: Cartago, Turrialba , 600m, 09º53'N 83º 38'W, January (F.T. Hovore)( USNM: ADP6384).

Diagnosis. Large, coppery beetles with apparently semi­catenate elytral interval and tridentate elytral apex. Male head tapered, female head rounded. Elytral foveae in interneurs 2,4, and 6 very large, adjacent intervals constricted at each fovea.

Description. Color and luster: Rufous. Dorsal surface bright metallic coppery green. Form: Head ( Fig. 25a ) behind eyes moderately elongate, tapered in male, rounded in female, occiput dimpled. Elytron ( Fig. 25 b) with apex markedly lobed medially, lateral tooth moderately small, sutural angle rounded, not projected acutely, but prolonged; foveae in interneurs 2,4, and 6 very large, adjacent intervals constricted at each fovea. Sternum VI (Fig, 25c) deeply V­notched in male, shallowly V­notched in female. Aedeagus ( Fig. 25 d) with ostium elongate, apex quadrate, longer than wide. Size: medium, 13.8 to 17.5mm in length, 3.4 to 4.6mm in width.

Other specimens examined. Costa Rica: 86 paratypes, 46 females, 40 males, from the following Conservation Areas: La Amistad Pacifico, Arenal Tilarán, Cordilleria Volcánica Central, Guanecaste, Tortuguero. Costa Rica, Alajuela, 1m, 790m, March, (H. J. Lezama)( MIUCR: ADP108575), 1m, 1f, 840m, March, (H. J. Lezama)( MIUCR: ADP108581, ADP108583), 1f, P.N. Volcán Tenorio, Upala, Bijagua, Albergue Heliconias, Sendero Heliconias, 700m, LN 422600,299100, April, (A. Lopez)( INBIO : INB0003476210), 1f, August, (A. Lopez)( INBIO : INB0003302213), 1m, 1f, November, (W. Porras)( INBIO : INB0003421011, INB0003421012), 1m, Upala Sector, Alto Los Brenes, 650m, LN 300600,422800, May, (A. Lopez)( INBIO : INB0003328227), 1f, Dos Rios, Finca San Gabriel, 600m, 85°22'W 10°50'N, May, (II curso Parataxonomos)( INBIO : CRI 000644731), 1f, Alajuela, Peñas Blancas, 84°37'W 10°04'N, March, (E. Cruz)( CMNC: ADP005548), 1f, April, (E. Cruz)( CMNC: ADP005552), 1m, September ­ October (E. Cruz)( CMNC: ADP005549), 1f, 1m, December (E. Cruz)( UASM: ADP005589, ADP005590), 1m, Sect. San Ramon de Dos Rios, Hda. Nueva Zelandia, 620m, LN 318100,381900, July, (D. Briceño)( INBIO : CRI 002468971), 1f, August, (F. A. Quesada )( INBIO : CRI 002483507), 1f, September, (C. Cano)( INBIO : CRI 002146268), 1m, 3f, Río San Lorencito, 850m, 84°35'W 10°13'N, LN 245500,470800, March, (A. Azofeila)( INBIO INB0003035066 INB0003035065), (R. A. Zuñiga )( INBIO : INB0003025841), (C. Viquez)( INBIO : INB0003025465), 1f, Sector Colonia Palmareña, 700m, LN 245900,475900, April, (E. Fletes)( INBIO : CRI 002248223); Cartago, 1m, 1250m, 83°46'22”W 9°43'02”N, August, (F. A. Quesada )( INBIO : CRI 000485925), 1f, Tucurrique, 83°43'W 9°51'N, ( MNHP: ADP001947), 1m, IICA EXPERIMENTAL STA­ TION ñ CATIE , Turrialba , Rio Reventazon, 600m, 83°39'W 9°53'N, LN 574947,208307, September, (G. Fonseca) ( INBIO : CRI 002093230), 1m, January, (F.T. Hovore)( FTHC: ADP006384), 1m, June, (E. Giesbert)( FSCA: ADP004425), 1f, (WAR: ADP004066), 1m, (A. Heyne)(WAR: ADP004060); Guanacaste, 1m, A.C.G. Liberia Mayorga, Estacíon Primario, 1000m, LN 322740,375198, March, (Espinoza)( INBIO : INB0003359189), 1m, Bagaces, Fortuna , Sendero Cabro Muco, 980m, LN 299151,410000, June, (A. Lopez)( INBIO : INB0003328390), 2f, 1m, Guanacaste, 9.0 km S Santa Cecilia, Estacíon Pitilla , 700m, 85°25'40” W 10°59'26”N, LN 330200,380200, January, (GNP Biodiversity Survey)( INBIO : CRI 001024198, CRI 001024200, CRI 001024208), 1f, (P. Rios)( INBIO : CRI 000605989), 1m, 2f, February, (P. Rios, C. Moraga, & R. Blanco)( INBIO : CRI 000175875, CRI 000175854, CRI 000124346), 1m, (P. Rios)( INBIO : CRI 001961467), 1m, March, (GNP Biodiversity Survey)( INBIO : CRI 000034318), 1f, (C. Moraga)( INBIO : CRI 002524455), 2m, (P. Rios, C. Moraga, & R. Blanco)( INBIO : CRI 000211525, CRI 000211528), 1f, 1m, April, (GNP Biodiversity Survey)( INBIO : CRI 000090738, CRI 001024199), 1m, (C. Moraga)( INBIO : CRI 000748636), 1f, April, (P. Knight)( INBIO : CRI 000336602), 1m, (C. Moraga)( INBIO : CRI 000637525), 1f, (P. Kazan, C. Moraga, & R. Blanco)( INBIO : CRI 000211526), 1m, (P. Rios)( INBIO : CRI 000615374), 1m, May, (F. Araya)( INBIO : CRI 000805112), 1m, (Z. Fuentes)( INBIO : CRI 000734037), 1m, (C. Moraga & P. Rios)( INBIO : CRI 000166083), 1f, 2m, (C. Moraga)( INBIO : CRI 000342062, CRI 000409528, CRI 002047454), 1f, (GNP Biodiversity Survey)( INBIO : CRI 001024207), 1f, 1m, (P. Opler)( INBIO : CRI 000959387, CRI 000389746), 1m, July, (GNP Biodiversity Survey)( INBIO : CRI 000129484), 1m, August, (C. Moraga)( INBIO : CRI 001869856), 2f, September, (C. Moraga & P. Rios)( INBIO : CRI 000609350), 1f, (P. Rios)( INBIO : CRI 002005180, CRI 000609311), 1m, November, (P. Opler)( INBIO : CRI 001355290), 1m, (C. Moraga)( INBIO : CRI 002018042), 1f, 1m, (P. Rios)( INBIO : CRI 000389747, CRI 000959394), 1m, December, (GNP Biodiversity Survey)( INBIO : CRI 001024209), 1f, (P. Opler)( INBIO : CRI 000610299), 1f, (C. Moraga)( INBIO : CRI 001975285), 1f, Volcán Cacao, Estacíon Cacao,1000 ñ 1400m, 85°25'40”W 10°59'26”N, LN 323300,375700, April, (C. Chaves)( INBIO : CRI 000633297); Heredia, 1f, 1m, 3.0 km S Puerto Viejo, Finca La Selva, Estacíon Biologica La Selva, 50 ñ 150m, 84°00'32”W 10°25'55”N, LN 535500,268000, January, (F.T. Hovore)( FTHC: ADP006447 ADP006448), 1f, January, (P.J. Landolt)( USNM: ADP004428), 2f, October, (Paul Hanson)( MIUCR: ADP 100517 ADP 100519), 1f, July, (H. A. Hespenheide )( HAH: ADP006915), 1f, April, ( INBIO : CRI 000627519), 1m, February, (H. A. Hespenheide )( HAH: ADP005091), 2f, April, (H. A. Hespenheide )( HAH: ADP005090 ADP005092); Limón, 1f, Cariari, Sector Cerro Cocori, Finca de E. Rojas, 150m, 83°42'29”W 10°35'29”N, LN 285700,568400, January, (E. Rojas) ( INBIO : CRI 001675492), 1f, Estacíon Hitoy Cerere, 100m, LN 643400,184600, January­December, (G. Carballo)( INBIO : INB0003430486), 1m, Estacíon Hitoy Cerere, Rio Cerere, Send. Bobocara, 740m, 83°01' W 9°40' N, LN 184750,639500, May, (W. Arana)( INBIO : INB0003114978); Puntarenas, 1f, 4.0 km S San Vito de Java, Coto Brus, Estacíon Biológica Las Alturas, 1520m, 83°50'17”W 8°47'07”N, LS 591100,322800, January, (M. A. Zumbado )( INBIO : CRI 000401758), 1m, Quepos, 80m, 84°09' W 9°24' N, LS 370900,448800, ( INBIO : CRI 001393998).

Specific epithet. The specific epithet, catbellae , is the Latinized genitive form of the combined name of the actress starring on the current TV program "JAG," Catherine Bell. These beetles share the forest with an elegant cat, the Jaguar.

Notes. This species has been collected more often than any other Agra in INBio's inventory process except the ubiquitous A. strangulata . An illustration of purpurea­group stylomere 2 is on the Agra purpurea web page.

Figures

FIGURE 25. A. catbellae, dorsal aspect: a) head, male, female; b) apex of elytron, male; c) Sternum VI, male (top), female (bottom), ventral aspect; d) aedeagus, dorsal (top), ventral (middle), left lateral (bottom) aspects. Measures (mm): HL = male 2.8, female 2.5; WE = 2.3; male SW = 4.2, female SW = 2.8; AL = 4.0.