Aspicera, Dahlbom, 1842

ROS-FARRÉ, P. & PUJADE-VILLAR, J., 2013, <strong> Revision of the genus <em> Aspicera </ em> Dahlbom, 1842 (Hym.: Figitidae: Aspicerinae) </ strong>, Zootaxa 3606 (1), pp. 1-110 : 8-12

publication ID	https://doi.org/ 10.11646/zootaxa.3606.1.1
publication LSID	lsid:zoobank.org:pub:3ED10277-CFD3-4D0D-B49A-C9F28AB6C955
DOI	https://doi.org/10.5281/zenodo.10539895
persistent identifier	https://treatment.plazi.org/id/03A24309-FFD3-614D-FF36-F940BADCFBDC
treatment provided by	Felipe
scientific name	Aspicera
status

Treatment

Redescription of Aspicera

Onychia Haliday in Walker, 1835: 162. Type-species: Evania ediogaster Rossi, 1790 , by monotypy. Homonym of Onychia Hübner, 1816: 334 (Lepidoptera) .

Aspicera Dahlbom, 1842: 6 . Type-species: Evania ediogaster Rossi, 1790 , by monotypy. Replacement name for Onychia Haliday by Reinhard, 1860: 242.

Bellona Giraud, 1860: 156 . Type-species: Evania ediogaster Rossi, 1790 , by monotypy. Unnecessary replacement name for Onychia Haliday. Homonym of Bellona Reichenbach, 1852 , (fossil bird).

Heteraspidia Belizin, 1952: 299–300 . Type-species: Heteraspidia foveata Belizin, 1952 , by monotypy. Synonymyzed by Ros-Farré & Pujade-Villar, 2011a: 54

Type species: Tenthredo scutellata Villers, 1789 (= Evania ediogaster Rossi, 1790 ; see taxonomic history of Aspicera )

Head densely pubescent on lower half, glabrous to sparsely pubescent on upper half. Lateral frontal carinae conspicuous, convergent towards antennal foramina ( Figs 1b & 1e View FIGURE 1 ) (in A. readae and A. elisendae they are parallel, straight with facial carinae ( Fig. 1c View FIGURE 1 ). Vertex straight or slightly incised. Ocelli weakly prominent to noticeably prominent. Occipital carinae conspicuous ( Figs 32a View FIGURE 32 & 44a View FIGURE 44 ). Occiput carinate ( Figs 2a View FIGURE 2 , 24b View FIGURE 24 & 34b View FIGURE 34 ) or without carinae ( Figs 7b View FIGURE 7 & 8b View FIGURE 8 ), usually coriaceous or alutaceous. Gena in lateral view expanded ( Fig. 44a View FIGURE 44 ) or not ( Fig. 15a View FIGURE 15 ) and curved ( Figs 32a View FIGURE 32 & 46a View FIGURE 46 ), although sometimes angled ( Fig. 27a View FIGURE 27 ), more or less carinate. Frons with variable sculpture, it can be coriaceous ( Fig. 1d View FIGURE 1 ), rugose or reticulate ( Figs 1c & 1f View FIGURE 1 ), sometimes also punctate ( Fig. 1e View FIGURE 1 ) or carinate ( Fig. 1a View FIGURE 1 ). Lateral frontal carinae conspicuous, sometimes weak near lateral ocelli. Antenna filiform or subclavate; female: 13-segmented (female of A. abbreviata has 14-segmented antennae), F1 narrower at basis than apex; male: 14-segmented, F1 sharply excavated ( Fig. 2b View FIGURE 2 ), with exception of A. scutellata and A. belizini , which have F1 very weakly modified ( Fig. 2a View FIGURE 2 ). Mesosoma sparsely pubescent. Lateral surface of pronotum: the microsculpture is coriaceous, except in A. abbreviata , which does not have microsculpture; the macrosculpture can be transverse carinae ( Figs 43a View FIGURE 43 & 44a View FIGURE 44 ), rugose ( Fig. 17a View FIGURE 17 ) or absent ( Figs 8a View FIGURE 8 & 30a View FIGURE 30 ). Pronotal plate conspicuous, subpronotal plate present, noticeably projected ahead in A. aculeata ( Fig. 1a View FIGURE 1 ). Scutum coriaceous, rarely shiny ( Fig. 15b View FIGURE 15 ), with ( Fig. 42b View FIGURE 42 ) or without ( Fig. 27b View FIGURE 27 ) transverse carinae, sometimes also punctate ( Fig. 11b View FIGURE 11 ). Anteroadmedian line sharp ( Fig. 14b View FIGURE 14 ) to very weak ( Fig. 8b View FIGURE 8 ), reaching 1/3 to 1/2 scutum length. Median ridge sharp or effaced ( Fig. 8b View FIGURE 8 ), reaching from anterior margin of scutum to median mesoscutal furrow, ( Fig. 14b View FIGURE 14 ). Notauli complete, coriaceous or shiny, sometimes with transverse carinae inside. Median mesoscutal furrow reaching, at least, basal 1/3 of scutum, coriaceous or shiny, sometimes with transverse carinae inside. Parascutal sulcus anteriorly ending in front of tegula ( Figs 8a View FIGURE 8 & 27a View FIGURE 27 ) or continuing to anterior end of notauli ( Figs 24a View FIGURE 24 & 44a View FIGURE 44 ). Area between notauli and end of median mesoscutal furrow flat ( Figs 15a View FIGURE 15 & 8a View FIGURE 8 ) or prominent ( Figs 24a View FIGURE 24 , 40a View FIGURE 40 & 45a View FIGURE 45 ). Mesopleuron smooth or coriaceous, with coarse transverse carinae or with a furrow on basal 1/3, with sharp sculpture below it. Scutellum with scutellar spine, which is variable in length, usually pointed ( Figs 23b View FIGURE 23 & 32b View FIGURE 32 ), sometimes slightly blunt ( Fig. 12b View FIGURE 12 & 15b View FIGURE 15 ) or near absent ( A. kerzhneri ). Scutellar disc with two large and conspicuous foveae, in lateral view flat ( Fig. 44a View FIGURE 44 ) or convex ( Fig. 15a View FIGURE 15 ), the end of scutellar disc can be in the same plane as scutellar spine ( Fig. 44a View FIGURE 44 ) or sloping down towards apex of scutellar spine ( Figs 39a View FIGURE 39 & 40a View FIGURE 40 ). Propodeum pubescent, propodeal carinae sharp, dorsally parallel, ventrally divergent ( Fig. 2e View FIGURE 2 ). Forewing with radial cell open, R1 present, short or long (reaching half the distance between base of R1 and wing margin), but never reaching margin of wing, R2 straight or curved, Rs+M absent or slightly impressed. Wing surface pubescence reduced or not. Marginal pubescence of wing present or absent. Metatibia always with a sharp longitudinal ridge on the posterior surface. Petiole wider than long, with conspicuous longitudinal costulae ( Fig. 2e View FIGURE 2 ); 3 rd abdominal tergum saddle-shaped and smooth ( Fig. 2d View FIGURE 2 ). 4 th abdominal tergum with marked punctuation, usually not reaching posterior margin of 3 rd Tergum.

Diagnosis. Aspicera can be distinguished from other genera of Aspicerinae ‘ sensu stricto ’ by the following characters (Ros-Farré, 2007):

- Anacharoides , Pujadella , Omalaspis and Callaspidia lack scutellar spine (present in Aspicera ).

- In Balna the scutellar spine is reduced to a blunt point, and the scutellum has only one deep central fovea; in Aspicera the scutellar spine can be short but it is never reduced to a blunt point and the scutellum has two foveae.

- In Paraspicera the scutellar spine is blunt, thick and wide, the scutum never lacks microsculpture and the median sulcus is always narrow; in Aspicera the scutellar spine is slender and pointed, rarely very short, but if so the scutum lacks microsculpture and is distinctly carinate transversely, and the median sulcus is wide.

- In Prosaspicera there is a conspicuous median sulcus beginning at lateral ocelli and ending at occiput, with parallel longitudinal carinae at sides ( Ros-Farré & Pujade-Villar 2007), and the compound eyes are surrounded by a sharp continuous carina that is the union of the frontal carinae, vertex lateral carinae, occipital carina and facial impression ( Ros-Farré & Pujade-Villar 2007); both characters are absent in Aspicera .

World key to the species of Aspicera

1. Scutellar spine almost absent, made up by a distal elongation of the circumscutellar carina ( Figs 2f View FIGURE 2 , 4b View FIGURE 4 )................. 2

- Scutellar spine slender and pointed, rarely very short ( Fig 13b View FIGURE 13 ).................................................. 3

2. Scutellar foveae shallow, at least 1/2 scutellum length and with two longitudinal carinae inside ( Fig. 2f View FIGURE 2 ). Occiput without carinae. Lateral surface of propodeum not carinate. Mesoscutum with some punctuations............ A. foveata ( Belizin, 1952)

- Scutellar foveae well defined, about 1/3 scutellum length and with three longitudinal carinae inside ( Fig 4b View FIGURE 4 ). Occiput with longitudinal carinae dorsally and transverse carinae ventrally. Lateral surface of propodeum carinate transversely. Mesoscutum with transverse carinae........................................................... A. kerzhneri ( Kovalev, 1974)

3. Scutellum irregularly rugose/carinate, without longitudinal carinae except for the interfoveal carina ( Fig. 5b View FIGURE 5 ). Scutellar spine short and slender. Occiput rugose and carinate ( Fig. 5b View FIGURE 5 ). Lateral line incomplete, ending before anterior margin of scutum ( Fig. 5a View FIGURE 5 ). Wing pubescence reduced, marginal pubescence absent.................................................... 4

- Scutellum variable, but if it has rugose carinae, these are always longitudinal-like. Scutellar spine variable in length and wideness. Occiput smooth, carinate or rugose. Wing pubescence present or absent...................................... 5

4. Mesoscutum with coarse coriaceous sculpture ( Fig. 5b View FIGURE 5 ). Scutellar foveae with scutellar pits ( Fig. 5b View FIGURE 5 ). Female scutellar spine between 0.38 to 0.46 times length of scutellar disc...................... A. singularica Ros-Farré & Pujade-Villar n. sp.

- Mesoscutum with fine coriaceous sculpture ( Fig. 6b View FIGURE 6 ). Scutellar foveae without scutellar pits ( Fig. 6b View FIGURE 6 ). Female scutellar spine 0.20 times length of scutellar disc................................................... A. clarimontis Kieffer, 1907

5. Scutum always clearly punctate, without transverse carinae ( Figs 7b View FIGURE 7 & 11b View FIGURE 11 ). Antero-admedian lines very thin or indistinct. Median ridge effaced. Scutellar spine always short, 0.3 to 0.5 times length of scutellar disc........................... 6

- Scutum sometimes punctate between notauli and lateral margins, with or without transversal carinae. Antero-admedian lines usually conspicuous. Median ridge well developed ( Figs 12b View FIGURE 12 & 15b View FIGURE 15 ). Scutellar spine variable in length................ 10

6. Scutellar foveae smooth, very large and wide. Scutellar disc coriaceous ( Fig. 7b View FIGURE 7 )..................................................................................................... A. kovalevi Ros-Farré & Pujade-Villar n. sp.

- Scutellar foveae with sharp to effaced longitudinal carinae, in A. jantonii it is nearly smooth, variable in size. Scutellar disc carinate longitudinally and emarginate ( Figs 8b View FIGURE 8 & 10b View FIGURE 10 ).......................................................... 7

7. Scutum with sparse punctuation, at least between notauli, distance between punctuations larger than the diameter of them. Scutellar foveae with very weak to effaced longitudinal carinae ( Fig. 8b View FIGURE 8 ).......................................... 8

- Scutum with abundant punctuation, distance between punctuations similar to the diameter of them. Scutellar foveae with sharp longitudinal carinae ( Figs 10b View FIGURE 10 & 11b View FIGURE 11 )...................................................................... 9

8. Scutellar spine triangular. Scutellar foveae shallow, with effaced longitudinal carinae, nearly smooth ( Fig. 8b View FIGURE 8 ). Occiput not clearly carinate.................................................................. A. jantonii Ros-Farré n. sp.

- Scutellar spine elongated. Scutellar foveae deep, very slightly striate longitudinally ( Fig. 9b View FIGURE 9 ). Occiput with thin but clear longitudinal carinae on dorsal half and with transverse carinae on ventral half..................... A. adelae Ros-Farré n. sp.

9. Scutellar foveae small and entirely carinate longitudinally ( Fig. 10b View FIGURE 10 ). Scutellar spine narrow ( Fig. 10b View FIGURE 10 )................................................................................. A. carlestolrai Ros-Farré & Pujade-Villar n. sp.

- Scutellar foveae large, longitudinally carinate centrally (near interfoveal carina), smooth laterally ( Fig. 11b View FIGURE 11 ). Scutellar spine triangular (wide anteriorly) ( Fig. 11b View FIGURE 11 )................................... A. gemmae Ros-Farré & Pujade-Villar n. sp.

10. Scutellum very sharply convex, gibbous. Scutellar spine robust and blunt. End of scutellar disc uniformly narrowing to end of scutellar spine, so it is difficult to distinguish the beginning of scutellar spine. Scutellar foveae distinctly wide, scutellar pits sharply deep. Median mesoscutal furrow smooth and shining. Lateral line complete, reaching anterior margin of scutum ( Fig. 12a & b View FIGURE 12 )......................................................................... A. enormis Belizin, 1952

- Scutellum flat, to boardly convex. Scutellar spine always pointed. End of scutellar disc different, the beginning of scutellar spine can always be distinguished. Character combination different............................................. 11

11. Scutellum convex, but sharply depressed on central area. Scutellar spine very short, 0.19 length of scutellar disc. Scutellar foveae wide, coriaceous anteriorly and slightly carinate posteriorly ( Fig. 13b View FIGURE 13 )............... A. minutispina Belizin, 1952

- Scutellar disc never depressed on central area. Scutellar spine longer. Scutellar foveae variable, but never conspicuously coriaceous anteriorly...................................................................................... 12

12. Scutellar disc without interfoveal carina, weakly rugose. Scutellar spine long, 1.25 length of scutellar disc, and robust ( Fig. 14b View FIGURE 14 ). Lateral face of pronotum with sharp transverse and sinuous carinae. Notauli conspicuously coriaceous and with abundant transverse carinae. Lateral line complete, reaching anterior margin of scutum. Area between notauli sharply prominent in lateral view ( Fig. 14a View FIGURE 14 ).............................................................. A. elisendae Ros-Farré n. sp.

- Scutellar disc with interfoveal carina (in A. daurica it is dificult to see), with variable sculpture. Scutellar spine shorter. Lateral face of pronotum with or without carinae. Notauli coriaceous or with transverse carinae, only in A. readae and A. sibirica with both characters toguether. Area between notauli flat or prominent............................................... 13

13. Scutum and scutellum shiny, without microsculpture, sometimes with macrosculpture ( Figs 15b View FIGURE 15 ). Scutellum always clearly convex. Scutellar pits very deep. Interfoveal carina effaced on scutellar disc.................. A. abbreviata Belizin, 1952

- Scutum always coriaceous, in A. magdae very weakly so ( Fig. 39b View FIGURE 39 ). Scutellum variable, flat to convex. Scutellar pits present or absent. Interfoveal carina present, except in A. daurica ....................................................... 14

14. Wing pubescence not reduced, marginal pubescence long, wing sharply smoked, with veins very conspicuously impressed (Rs+M present). Occiput with thin but noticeable carinae. Lateral surface of pronotum transversely carinate dorsally ( Fig. 16a View FIGURE 16 ). Scutellar spine thin and slender. Scutellar disc with prominent interfoveal carina. Scutellar foveae with very weak to sharp transverse carinae anteriorly and longitudinal carinae posteriorly ( Fig. 16b View FIGURE 16 )............... A. sergioi Ros-Farré n. sp.

- Wing pubescence variable. Occiput variable, with or without carinae (but if it has noticeable carinae then lateral surface of pronotum is clearly rugose or with sharp transverse carinae). Lateral surface of pronotum coriaceous to carinate. Interfoveal carina variable. Scutellar foveae variable........................................................................ 15 15. Scutellar foveae with, at least, two sharp longitudinal carinae that continue, together with interfoveal carina, along scutellar disc. Scutellar pits sharply deep. Scutum with sharp coriaceous sculpture ( Figs 17b View FIGURE 17 & 18b View FIGURE 18 )......................... 16

- Scutellar foveae different, they can have weak longitudinal carinae except for A. daurica , which has sharp longitudinal carinae, but in this species interfoveal carina is effaced on scutellar disc ( Fig. 38b View FIGURE 38 ). Scutellar pits present or absent. Scutum variable.................................................................................................... 17

16. Scutum with transverse carinae ( Fig. 17b View FIGURE 17 )............................................. A. hartigi Dalla Torre, 1889

- Scutum coriaceous ( Fig. 18b View FIGURE 18 )....................................... A. buffingtoni Ros-Farré & Pujade-Villar n. sp.

17. Scutellar disc with prominent interfoveal carina, (if there are lateral carinae, they are not so prominent). Carinae of occiput medium impressed ( Fig. 19b View FIGURE 19 ). Lateral line complete ( Fig. 19a View FIGURE 19 ). Marginal pubescence of wing long, exremely long in the dorsal margin between basis of wing and radial cell ( Fig. 3a View FIGURE 3 )..................................... A. tomasi Ros-Farré n. sp.

- Interfoveal carina variable. Marginal pubescence of wing different, never so long in dorsal margin. Character combination different.............................................................................................. 18

18. Occiput with sharp carinae ( Fig. 2c View FIGURE 2 ). Lateral surface of pronotum coriaceous. Scutellum very slightly convex, nearly flat. Marginal pubescence of wing absent. Scutellar disc with a medium prominent interfoveal carina, which continues straight until half of length, and undulate posteriorly.............................................. A. utahensis Ashmead, 1896

- If occiput has sharp carinae, lateral surface of pronotum is clearly rugose or with sharp transversal carinae. Scutellum flat to convex. Marginal pubescence of wing variable. Scutellar disc with weak to prominent interfoveal carina............... 19

19. Occiput with or without weak carinae ( Figs 23b View FIGURE 23 , 25b View FIGURE 25 & 30b View FIGURE 30 ). Lateral surface of pronotum coriaceous, rarely with scarce and weak transverse carinae on dorsal half ( Figs 23a View FIGURE 23 , 25a View FIGURE 25 & 30a View FIGURE 30 ). Scutellar disc with a prominent interfoveal carina; lateral carinae of scutellar disc, if present, not so prominent as interfoveal carina ( Figs 23a View FIGURE 23 , 25a View FIGURE 25 , 31a & 31b View FIGURE 31 )........................ 20

- Occiput with sharp to very sharp carinae ( Figs 39b View FIGURE 39 & 40b View FIGURE 40 ). Lateral surface of pronotum clearly rugose or with sharp transverse carinae ( Figs 40a View FIGURE 40 , 42a View FIGURE 42 & 48a View FIGURE 48 ). Scutellar disc only with a noticeable, but not prominent, interfoveal carina; lateral carinae of scutellar disc are similar or weaker than interfoveal carina ( Figs 40a View FIGURE 40 , 42a & b View FIGURE 42 ).................................... 34

20. Lateral line complete, reaching anterior margin of scutum ( Figs 24a & b View FIGURE 24 ). Marginal pubescence of wing long............ 21

- Lateral line incomplete, ending before anterior margin of scutum ( Figs 25a & b View FIGURE 25 ). Marginal pubescence of wing variable... 24

21. Lateral surface of pronotum without transversal carinae ( Fig. 21a View FIGURE 21 ). Scutellar foveae with weak longitudinal carinae posteriorly ( Fig. 21b View FIGURE 21 ). Mesopleuron coarsely coriaceous on anterior 1/3................................. A. sibirica Kieffer, 1901

- Lateral surface of pronotum with a few weak transverse striae or carinae. Scutellar foveae smooth. Mesopleuron with clear punctate sculpture or with some rugose carinae on anterior 1/3................................................. 22

22. Scutellar spine short, 0.5 length of scutellar disc. Scutellar foveae sharply wide and shallow ( Fig. 22c View FIGURE 22 ). Notauli transversely carinate and weakly alutaceous ( Fig. 22b View FIGURE 22 ). Mesopleuron with clear punctate sculpture on anterior 1/3.................................................................................................... A. attenuata Belizin, 1952

- Scutellar spine long, 0.7 to 0.9 length of scutellar disc ( Figs 23b View FIGURE 23 & 24b View FIGURE 24 ). Scutellar foveae variable. Notauli coriaceous or alutaceous. ( Figs 23b View FIGURE 23 & 24b View FIGURE 24 ). Mesopleuron with some rugose carinae on anterior 1/3................................... 23

23. Notauli coriaceous. Median mesoscutal furrow wide anteriorly. Occiput coriaceous, with carinae ( Fig. 24b View FIGURE 24 ). Area between notauli prominent in lateral view ( Fig. 24a View FIGURE 24 ), pointed centrally. Scutellar disc rectangular, seen in lateral view at the same level than scutellar spine, which is flat........................................ A. martae Ros-Farré & Pujade-Villar n. sp.

- Notauli smooth centrally, alutaceous near margins. Median mesoscutal furrow narrow dorsally. Occiput rugose behind ocelli, withouth carinae ( Fig. 23b View FIGURE 23 ). Area between notauli not prominent in lateral view, curved. Scutellar disc oval, slightly sloping posteriorly towards basis of scutellar spine, which is directed downwards ( Fig. 23a View FIGURE 23 )..................................................................................................... A. annae Ros-Farré & Pujade-Villar n. sp.

24. Scutellar disc very flat and wide, smoothly coriaceous and weakly rugose; lateral bar narrow and complete, scutelar disc entirely and sharply Marginated. Scutellar foveae very large. Scutum coriaceous. Median ridge weak near anterior margin of median mesoscutal furrow ( Fig. 25b View FIGURE 25 ). Mesopleuron sharply coriaceous posterodorsally............................................................................................... A. marginata Ros-Farré & Pujade-Villar n. sp.

- Scutellar disc and lateral barr different; sometimes marginate, rarely sharply or entirely, lateral bar never so narrow. Scutellar foveae usually smaller. Scutum variable. Median ridge different. Mesopleuron variable............................ 25

25. Scutellar foveae with one or two thin oblique carinae on each side of interfoveal carina. Scutellar spine extremely short. Notauli and median mesoscutal furrow transversely striate ( Fig. 26b View FIGURE 26 )....... A. santamariai Ros-Farré & Pujade-Villar n. sp.

- Scutellar foveae different. Character combination different.................................................... 26

26. Scutellar foveae rounded, transversely carinate anteriorly and rugose posteriorly, deep and small, lateral bar (between scutellar fovea and scutellum margin) very wide. Scutellar spine short, equilateral-triangle-shaped. ( Fig. 27b View FIGURE 27 ). Ocelli clearly prominent................................................................... A. mireiae Ros-Farré & Pujade-Villar n. sp.

- Scutellar foveae different, lateral bar not as wide as in A. mireiae . Scutellar spine usually longer than wide ( Figs 28b View FIGURE 28 & 29b View FIGURE 29 ). Ocelli slightly prominent............................................................................... 27

27. Scutellar disc not swollen posterior to scutellar foveae ( Figs 28a View FIGURE 28 , 29a View FIGURE 29 & 31a View FIGURE 31 )...................................... 28

- Scutellum swollen posterior to scutellar foveae (35a), sometimes very slightly so, in this case marginal pubescence of wing very noticeable...................................................................................... 31

28. Lateral frontal carinae absent on dorsal half, near ocelli, in this area there is a clear and abundant punctate sculpture ( Fig. 1e View FIGURE 1 )............................................................... A. punctifrons Ros-Farré & Pujade-Villar n. sp.

- Lateral frontal carinae complete, punctuations never so sharp ( Figs 1f & 1d View FIGURE 1 )...................................... 29

29. Scutellar spine short and wide. Scutum slightly punctate and coriaceous ( Fig. 29b View FIGURE 29 )............. A. teresae Ros-Farré n. sp.

- Scutellar spine long and thin. Scutum coriaceous, with abundant to scarce transverse carinae. ( Figs 30b View FIGURE 30 & 31b View FIGURE 31 ).......... 30

30. Scutellum sloping downwards on each side of interfoveal carina (roof-shaped), which is extremely prominent and sharp. Scutellar disc sharply rugose and with some sinuous longitudinal carinae ( Fig. 30b View FIGURE 30 )................................................................................................... A. carinata Ros-Farré & Pujade-Villar n. sp.

- Scutellum flat on each side of interfoveal carina, which is prominent but fine. Scutellar disc rugose, without longitudinal carinae after foveae ( Fig. 31b View FIGURE 31 )........................................................... A. dianae Ros-Farré n. sp.

31. Marginal pubescence of wing present, uniformly distributed in all the wing margin................................. 32

- Marginal pubescence of wing absent or discontinuous........................................................ 33

32. Notauli narrow. Scutellum with sharp longitudinal carinae. Scutellar foveae coriaceous on posterior 1/3 ( Fig. 32b View FIGURE 32 ).............................................................................. A. julii Ros-Farré & Pujade-Villar n. sp.

- Notauli wide. Scutellum with some fine longitudinal carinae. Scutellar foveae with weak carinae ( Figs 33a & b View FIGURE 33 )............................................................................................ A. proxima Belizin, 1952

33. Scutum very coriaceous. Scutellar foveae shallow, with abundant transverse carinae on anterior half and scarce longitudinal carinae on posterior half ( Fig. 34b View FIGURE 34 )...................................... A. kiefferi Ros-Farré & Pujade-Villar n. sp.

- Scutum with short transverse carinae, mostly between notauli. Scutellar foveae deep and rugose ( Fig. 35b View FIGURE 35 )................................................................................ A. zuparcoi Ros-Farré & Pujade-Villar n. sp.

34. Scutellar disc convex after foveae ( Figs 37a View FIGURE 37 , 38b View FIGURE 38 & 39 View FIGURE 39 ) Marginal pubescence of wing variable....................... 35

- Scutellar disc flat after foveae ( Figs 44a View FIGURE 44 & 45b View FIGURE 45 ). Marginal pubescence of wing starting posterior to R2, conspicuous...... 42

35. Lateral line complete, reaching anterior margin of scutum. Wing pubescence not reduced, marginal pubescence long and dense. Scutum without transverse carinae.................................................................. 36

- Character combination different.......................................................................... 37

36. Scutellar foveae weakly sculptured and shallow ( Fig. 36b View FIGURE 36 ). Scutellar disc sharply convex.......... A. porif Ros-Farré n. sp.

- Scutellar foveae smooth and deep ( Fig. 37b View FIGURE 37 ). Scutellar disc weakly convex................. A. simulacrum Belizin, 1952

37. Punctuations of 4 th metasomal segment reaching posterior margin of 3 rd segment in dorsal view. Interfoveal carina not continuing on scutellar disc. Scutum with scarce microsculpture ( Fig. 38b View FIGURE 38 ). Occiput with transverse carinae, even behind ocelli. Wing membrane hyaline.................................................................. A. daurica Belizin, 1952

- Punctuations of 4 th metasomal segment not reaching posterior margin of 3 rd segment in dorsal view. Interfoveal carina present on scutellar disc ( Fig. 39b View FIGURE 39 ). Character combination different................................................... 38

38. Lateral line incomplete, only reaching basal 2/3 of scutum ( Fig. 39b View FIGURE 39 & 40a View FIGURE 40 )...................................... 39

- Lateral line complete, reaching anterior margin of scutum ( Figs 43a & b View FIGURE 43 ); in A. sculpturata , although they are very conspicuous they end just before anterior margin of scutum ( Fig. 42a View FIGURE 42 ).................................................. 40

39. Scutellar foveae with weak oblique carinae, scutellar pits deep. Scutum very slightly coriaceous. Occiput with short longitudinal carinae behind ocelli ( Fig. 39b View FIGURE 39 ). Lateral surface of pronotum entirely carinate transversely ( Fig. 39a View FIGURE 39 )................................................................................... A. magdae Ros-Farré & Pujade-Villar n. sp

- Scutellar foveae smooth, scutellar pits absent. Scutum with noticeable coriaceous sculpture. Occiput with long longitudinal carinae behind ocelli ( Fig. 40b View FIGURE 40 ). Lateral surface of pronotum transversely carinate dorsally and ventrally, coriaceous centrally ( Fig. 40a View FIGURE 40 ).......................................................... A. belizini Ros-Farré & Pujade-Villar n. sp.

40. Scutum with noticeable rugose carinae ( Fig. 42b View FIGURE 42 ). Notauli sharply carinate transversely............................ 41

- Scutum usually coriaceous, but sometimes with weak and short transverse carinae ( Fig. 41b View FIGURE 41 ). Notauli weakly carinate transversely................................................................ A. suecica Dalla Torre & Kieffer, 1910

41. Scutellar pits deep, scutellum rugose ( Fig. 42b View FIGURE 42 ). Lateral line nearly reaching anterior margin of scutum ( Fig. 42a View FIGURE 42 ). ......................................................................................... A. scutellata ( Villers, 1789)

- Scutellar pits absent, scutellum with parallel longitudinal carinae ( Fig. 43b View FIGURE 43 ). Lateral line complete, reaching anterior margin of scutum ( Fig. 43a View FIGURE 43 )................................................................. A. blancae Ros-Farré n. sp.

42. Subpronotal plate conspicuous and projected ahead ( Fig. 44b View FIGURE 44 ). Frontal and lateral frontal carina very conspicuous ( Fig. 1a View FIGURE 1 ). Scutellar spine extremely robust, wide and long, scutellar foveae noticeably rugose, scutellar pits sharply deep ( Fig. 44b View FIGURE 44 )................................................................... A. forshzarai Pujade-Villar & Ros-Farré n. sp.

- Subpronotal plate not projected or only projected basally ( Fig. 45b View FIGURE 45 ). Lateral frontal carinae variable, frontal carinae absent or medium impressed ( Fig. 1c View FIGURE 1 ). Scutellar spine never so robust, wide and long, scutellar foveae smooth, with weak longitudinal carinae or slightly alutaceous, scutellar pits absent ( Figs 45b View FIGURE 45 , 46b View FIGURE 46 & 48b View FIGURE 48 )........................................ 43

43. Lateral frontal carinae, parallel, straight with facial carinae, which are also parallel ( Fig. 1c View FIGURE 1 ). Notauli coriaceous with scarce transverse carinae ( Fig. 45b View FIGURE 45 )........................................... A. readae Ros-Farré & Pujade-Villar n. sp.

- Lateral frontal carinae, confluent towards antennal foramina, not straight with facial carinae, which are divergent ( Fig. 1b View FIGURE 1 ). Notauli shiny, with transverse carinae..................................................................... 44

44. Scutellar spine robust, wide basally, medium length (0.55 to 0.65 length of scutellar disc). Scutellar disc with sharp, prominent and sinuous longitudinal carinae ( Fig. 46b View FIGURE 46 )............................................ A. caminali Ros-Farré n. sp.

- Scutellar spine slender, narrow basally, long (0.65 to 1.20 length of scutellar disc). Scutellar disc different ( Fig. 47b View FIGURE 47 , 48b View FIGURE 48 & 49b View FIGURE 49 )................................................................................................ 45

45. Scutellar foveae shallow, rugose, scutellar disc rugose ( Fig. 47b View FIGURE 47 ).............. A. robusta Ros-Farré & Pujade-Villar n. sp.

- Scutellar foveae deeper, smooth or with effaced sculpture, scutellar disc with straight longitudinal carinae ( Figs 48b View FIGURE 48 & 49b View FIGURE 49 ). (species difficult to separate)............................................................................ 46

46. Robust insect (males>3.0mm., females> 3.4mm.). Scutum with or without weak transverse carinae, notauli and median mesoscutal furrow with small and weak transverse carinae ( Fig. 48b View FIGURE 48 ). Interfoveal carina continuing along scutellar disc to beginning of scutellar spine................................................................. A. longispina Kieffer, 1901

- Insect not robust (males<2.7mm., females unknown). Scutum with sharp transverse carinae, notauli and median mesoscutal furrow with sharp transverse carinae ( Fig. 49b View FIGURE 49 ). Interfoveal carina continuing along scutellar disc and spine until last 1/3 of scutellar spine................................................... A. danielssoni Ros-Farré & Pujade-Villar n. sp.