Pupopsis subtorquilla, Wu, Min & Gao, Linhui, 2010

Wu, Min & Gao, Linhui, 2010, A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China, Zootaxa 2725, pp. 1-27 : 20

11. Pupopsis subtorquilla sp. nov.

Figs. 1 , 3 I, 11A–11C; Table 2

Type material. Holotype: HBUMM-05429–specimen 2 (fms), Yuxushan Hill, 10 km north of town of Wenxian, site No. 0 45, 1070 m a.s.l., 32°57ʹ35.5ʹ N, 104°40ʹ41.6ʹ E, Wenxian County, S. Gansu, China, 2006–IX–27, leg. M. Wu, J. Liu, W. Zheng & L. Gao. Paratypes: HBUMM-05429–specimens 1, 3–13, (12 fms), same collection data as holotype; HBUMM-05545–specimens 1–3, (3 fms), 829 m a.s.l., 32°51ʹ44.7ʹ N, 104°50ʹ48.2ʹ E, Hendan Village, along national highway no. 212, S. Gansu, China, 2006–IX–29, leg. M. Wu, J. Liu, W. Zheng & L. Gao; HBUMM- 0 0 349 (1 fms), Bikou, 720 m a.s.l., 32°48ʹ, 105°12ʹE, Wenxian County, Gansu, China, 2004–IV–18, leg. M. Wu; HBUMM-00314–specimens 1–4 (4 fms), Wenxian, Shangdanpu, 1450 m a.s.l., 33°0ʹN, 104°36ʹE, 2004–IV–22, leg. M. Wu; HBUMM-00337–specimens 1–3 (3 fms), Wenxian, 2004–IV–7, leg. M. Wu.

Type locality. Yuxushan Hill, 10 km north of Wenxian, Wenxian County, S. Gansu.

Material examined. Type material.

Distribution. Wenxian County, Gansu.

Etymology. The new species is named for its congener, P. torquilla , to which it is taxonomically closest. The epithet is a noun in apposition.

Diagnosis. Shell with 6.500–7.500 whorls. Height 7.0–8.4, diam. maj. 3.5–4.5. Parietal tooth present. Columellar tooth prominent and expanded inward. Palatal tooth ridge-like.

Shell. Bulbous-ovate; apex not acuminate; dextral; thick-shelled; solid; opaque; glossy; with 6.500–7.025– 7.500 whorls. Whorls never spirally grooved; somewhat convex. Embryonic shell smooth; not polished; with 1.250–1.554–2.500 whorls. Postnuclear whorls wrinkled. Suture simple. Last whorl gradually ascending toward aperture; rounded at periphery. Shell 7.0–7.8–8.4 high; 3.5–4.0– 4.5 in diam. maj. Height/diam. maj. ratio 1.79– 1.97–2.30. Aperture subovate; almost vertical; its insertions separated; 2.5–2.9–3.4 high, 2.3–2.6–2.8 wide. Ratio of shell height to aperture height 2.27–2.71–3.16. Palatal margin rounded. Peristome expanded, sharp. Palatal tooth ridge-like, extending through nearly the entire body whorl. Parietal callus distinct. Angular tubercle and parietal tooth present. Columellar margin reflexed; with one tooth which is hardly visible in apertural view but extends inward along the columella. Umbilicus a narrow slit. Shell, including upper whorls, uniformly light corneousbrown; aperture white.

Remarks. The new species is close to P. torquilla due to similar size and the development of the apertural armature. However, the shell of the new species is much more swollen, with much fewer whorls and the columellar tooth is almost invisible in apertural view.

Figures

FIGURE 1. Distribution map of the genus Pupopsis. Red dot, Pupopsis dissociabilis (Sturany, 1900); blue dot, P. retrodens (Martens, 1879); blue square, P. maoxian sp. nov.; red square, P. rhodostoma sp. nov.; green dot, P. subpupopsis sp. nov.; green square, P. yengiawat sp. nov. Two grey dots each with two or five species as indicated in the boxes on the map. The species P. paraplesius (Sturany, 1900), P. polystreptus (Sturany, 1900) and P. zilchi sp. nov. are not mapped due to no precise locality information available.

FIGURE 3. Apertural view of Pupopsis species. A, Pupopsis pupopsis (Gredler, 1898), SMF- 42076; B, Pupopsis retrodens (Martens, 1879), HBUMM- 06460 – specimen 1; C-D, Pupopsis torquilla (Möllendorff, 1901). C, lectotype, SMF- 42082; D, HBUMM- 05584 – specimen 1; E, Pupopsis hendan sp. nov., holotype, HBUMM- 05592 – specimen 2; F, Pupopsis maoxian sp. nov., holotype, HBUMM- 01023 – specimen 1; G, Pupopsis rhodostoma sp. nov., holotype, HBUMM- 05872 – specimen 2; H, Pupopsis subpupopsis sp. nov., holotype, HBUMM- 05676 – specimen 1; I, Pupopsis subtorquilla sp. nov. A, holotype, HBUMM- 05429 – specimen 2; J, Pupopsis yengiawat sp. nov., holotype, HBUMM- 04439 – specimen 1; K, Pupopsis yuxu sp. nov., Holotype, HBUMM- 06089 – specimen 3; L, Pupopsis zilchi sp. nov., holotype, SMF- 334777.