Pupopsis maoxian, Wu, Min & Gao, Linhui, 2010

Wu, Min & Gao, Linhui, 2010, A review of the genus Pupopsis Gredler, 1898 (Gastropoda: Stylommatophora: Enidae), with the descriptions of eight new species from China, Zootaxa 2725, pp. 1-27 : 17

8. Pupopsis maoxian sp. nov.

Figs. 1 , 3 F, 8E; Table 2

Type material. Holotype: HBUMM-01023–specimen 1 (fms); 5 km from Fengyi Town on the road to Wenchuan County, 1574 m a.s.l., 31°39ʹ35.4ʹ N, 103°48ʹ48.7ʹ E, Maoxian County, NW Sichuan, China, 2004–X–11, leg. M. Wu. Paratypes: HBUMM-01023–specimen 2 (fms), same data as holotype; HBUMM-01794–specimen 1 (broken fms), Hongqi Hill (ca. 2 km south of the town of Maoxian ), Maoxian County, NW Sichuan, China, 2004–IX, leg. M. Wu.

Type locality. 5 km from Fengyi Town on the road to Wenchuan County, Maoxian County, NW Sichuan.

Material examined. Type material.

Distribution. Sichuan ( Maoxian County).

Etymology. The new species is named after the county name “ Maoxian .” The epithet is a noun in apposition.

Diagnosis. Shell with 8.125 whorls. Height 8.8–8.9, diam. maj. 2.2–3.4. Parietal tooth absent. Columellar tooth prominent and expanded inward. Palatal wall with a ridge-like tooth.

Shell. Fusiform; apex not acuminate; dextral; thin-shelled; solid; semitranslucent; glossy; with 8.125–8.125– 8.125 whorls. Whorls never spirally grooved; convex. Embryonic shell smooth; polished; with 1.500–1.563–1.625 whorls. Postnuclear whorls wrinkled. Suture simple. Last whorl gradually ascending toward aperture; rounded at periphery. Shell 8.8–8.8–8.9 high; 2.2–2.8– 3.4 in diam. maj. Height/diam. maj. ratio 2.64–3.36–4.08. Aperture ovate; slightly oblique; its insertions separated; 1.8–2.3–2.8 high, 2.0–2.1–2.1 wide. Ratio of shell height to aperture height 3.17–3.97–4.77. Peristome expanded; sharp. Palatal margin rounded. Palatal tooth present, ridgelike, extending approximately 1.25 whorls inward. Parietal callus indistinct; with angular tubercle. Parietal tooth absent. Columellar margin reflexed; with one prominent tooth which is expanded inward. Umbilicus a narrow slit. Shell uniformly light brownish-white; aperture white.

Remarks. This new species is grouped into the genus Pupopsis because of its typical apertural armature. The unique shell contour with a pointed apex and a narrowed body whorl, and the combination of the absence of a parietal tooth and the presence of both palatal, ridge-like tooth and columellar tooth set P. m a o x i a n apart from all other species in the genus.

This new species lives in a habitat charaterised by seasonal change of aridity and wetness. The detailed environmental characters are almost the same as described in Wu et al. (2003) for Cathaica (Pliocathaica) radiata Pilsbry, 1934 .

Figures

FIGURE 1. Distribution map of the genus Pupopsis. Red dot, Pupopsis dissociabilis (Sturany, 1900); blue dot, P. retrodens (Martens, 1879); blue square, P. maoxian sp. nov.; red square, P. rhodostoma sp. nov.; green dot, P. subpupopsis sp. nov.; green square, P. yengiawat sp. nov. Two grey dots each with two or five species as indicated in the boxes on the map. The species P. paraplesius (Sturany, 1900), P. polystreptus (Sturany, 1900) and P. zilchi sp. nov. are not mapped due to no precise locality information available.

FIGURE 3. Apertural view of Pupopsis species. A, Pupopsis pupopsis (Gredler, 1898), SMF- 42076; B, Pupopsis retrodens (Martens, 1879), HBUMM- 06460 – specimen 1; C-D, Pupopsis torquilla (Möllendorff, 1901). C, lectotype, SMF- 42082; D, HBUMM- 05584 – specimen 1; E, Pupopsis hendan sp. nov., holotype, HBUMM- 05592 – specimen 2; F, Pupopsis maoxian sp. nov., holotype, HBUMM- 01023 – specimen 1; G, Pupopsis rhodostoma sp. nov., holotype, HBUMM- 05872 – specimen 2; H, Pupopsis subpupopsis sp. nov., holotype, HBUMM- 05676 – specimen 1; I, Pupopsis subtorquilla sp. nov. A, holotype, HBUMM- 05429 – specimen 2; J, Pupopsis yengiawat sp. nov., holotype, HBUMM- 04439 – specimen 1; K, Pupopsis yuxu sp. nov., Holotype, HBUMM- 06089 – specimen 3; L, Pupopsis zilchi sp. nov., holotype, SMF- 334777.