Eviulisoma cetafi, Enghoff, 2018

Enghoff, Henrik, 2018, A mountain of millipedes VII: The genus Eviulisoma Silvestri, 1910, in the Udzungwa Mountains, Tanzania, and related species from other Eastern Arc Mountains. With notes on Eoseviulisoma Brolemann, 1920, and Suohelisoma Hoffman, 1963 (Diplopoda, Polydesmida, Paradoxosomatidae), European Journal of Taxonomy 445, pp. 1-90 : 28-30

Eviulisoma cetafi sp. nov.

urn:lsid:zoobank.org:act: 2754 E233-F5D6-493B-B410- 1067 FF17 A 521

Figs 7B, D , 13

Diagnosis

Differs from other species of the E. kwabuniense group by having the solenophore two-lobed, its dorsal lobe developed as a very strong hook, hook much larger than those seen in certain other species ( E. ejti sp. nov., E.akkariae sp. nov., E. nessiteras sp. nov.).

Etymology

The species name honours CETAF, Consortium of European Taxonomic Facilities, www.cetaf.org, in recognition of the immense importance of CETAF for natural history collections in Europe and for collections-based research.

Material studied (total: 12 ♂♂)

Holotype

TANZANIA: ♂, Iringa Region, Udzungwa Scarp FR, above Chita Village , 1300–1400 m a.s.l., 26 Oct.– 14 Nov. 1984, pitfall traps in montane rain forest, N. Scharff leg. ( ZMUC).

Paratypes

TANZANIA: 2 ♂♂, Iringa Region, Udzungwa Scarp FR, above Chita Village , 1450 m a.s.l., 4–9 Nov. 1984, pitfall traps in montane rain forest, N. Scharff leg. ( ZMUC) ; 4 ♂♂, Iringa Region, Udzungwa Scarp FR, above Chita Village , 1500 m a.s.l., 2–13 Nov. 1984, pitfall traps in montane rain forest, N. Scharff leg. ( ZMUC) ; 2 ♂♂, Iringa Region, Udzungwa Scarp FR, above Chita Village , 1500–1550 m a.s.l., 2–13 Nov. 1984, pitfall traps in montane rain forest, N. Scharff leg. ( ZMUC) ; 1 ♂, Iringa Region, Udzungwa Scarp FR, above Chita Village , 1300 m a.s.l., 2–6 Nov. 1984, pitfall traps in montane rain forest, N. Scharff leg. ( ZMUC) ; 2 ♂♂, Iringa Region, Udzungwa Scarp FR, above Chita Village , 1400 m a.s.l., 4–5 Nov. 1984, pitfall traps in montane rain forest, N. Scharff leg. ( ZMUC) .

Description (male)

SIZE. Length 20 mm, max. width 1.9–2.1 mm.

COLOUR. After 33 years in alcohol all whitish.

ANTENNAE. Reaching back to middle of ring 3.

BODY RINGS. Paranota indicated by very faint ridges on body ring 2 (as in Fig. 4C ), otherwise completely absent. Stricture between pro- and metazonite not striolate. A transverse row of setae on all body rings, but the majority of setae abraded in most specimens.

HYPOPROCT. Large, trapezoid, with three prominent apical tubercles, middle tubercle larger.

LEGS. Length 1.2× body width. Relative lengths of podomeres: femur> prefemur> tarsus> tibia> postfemur. Scopulae ( Fig. 7B, D ) on femur, postfemur, tibia and tarsus until ca midbody, thereafter gradually disappearing.

STERNUM 5. A small trapezoid process between legs 4.

STERNUM 6. Deeply excavated. Rim of excavation simple.

GONOPODS ( Fig. 13 ). Coxal lobe (cxl) moderate. Prefemoral part (prf) ca half as long as acropodite. Mesal acropodital process (map) a long, smooth, almost straight rod, apically pointed, subapically with small triangular side branch. Intermediate acropodital process (iap) a very slender, straight rod, densely covered in long spines on part of its surface, especially on apical part (Fig. E). Solenophore (sph) deeply split into two long lobes; dorsal lobe (sph-d) a very large, strong hook, almost reaching to tip of acropodital processes; ventral lobe (sph-v) much shorter, lanceolate; no intermediate process between sph-d and sph-v; internal surface of sph with a ridged area (ra, Fig. 13D, F ).

Distribution and habitat

Known only from the Udzungwa Scarp FR. Altitudinal range 1300–1550 m a.s.l. Habitat: montane rain forest. Collected together with E. chitense sp. nov.

Figures

Fig. 4. Eviulisoma spp. A–B. E. paradisiacum sp. nov., ♂. A. Head and rings 1–8, lateral view. B. Head and collum, frontal view. C. E. Zebra sp. nov., ♂, head and rings 1–3, lateral view. Abbreviation: pn = paranotum. Scale bars: 0.2 mm.

Fig. 7. Eviulisoma spp., scopulae on postfemur (pof), femur (fe), tibia (ti) and tarsus (ta) of male legs. A. E. paradisiacum sp. nov., paratype. B, D. E. cetafi sp. nov., paratype. C. E. ejti sp. nov., paratype. E–F. E. coxale sp. nov., paratype from Mito Mitatu. Scale bars: A, C, E = 0.05 mm; B, F = 0.01 mm; D = 0.1 mm.

Fig. 13. Eviulisoma cetafi sp. nov., paratype, right gonopod. A. Mesal view. B. Meso-dorsal view. C. Latero-ventral view. D. Ventral view. E. Spiny tip of iap. F. Ridged area of solenophore with tip of solenomere sticking out. Abbreviations: cxl = coxal lobe; iap = intermediate acropodital process; map = mesal acropodital process; prf = prefemoral part; ra = ridged area of solenomere; slm = solenomere; sph-d, sph-i, sph-v = dorsal, intermediate and ventral lobes of solenophore. Scale bars: A–D = 0.1 mm; E = 0.01 mm; F = 0.02 mm.