Athleta (Volutospina) noetlingi ( Cossmann & Pissarro, 1909 )

Merle, Didier, Pacaud, Jean-Michel, Métais, Grégoire, Bartolini, Annachiara, Lashari, Rafiq A., Brohi, Imdad A., Solangi, Sarfraz H., Marivaux, Laurent & Welcomme, Jean-Loup, 2014, Volutidae (Mollusca: Gastropoda) of the Lakhra Formation (Earliest Eocene, Sindh, Pakistan): systematics, biostratigraphy and paleobiogeography, Zootaxa 3826 (1), pp. 101-138 : 118

Athleta (Volutospina) noetlingi ( Cossmann & Pissarro, 1909)

Volutospina noetlingi Cossmann & Pissarro, 1909: 24 , pl. 2, figs 23–24.

? Voluta cithara (non Voluta cithara Lamarck, 1803 ). d’Archiac & Haime, 1853: 325, pl. 32, figs 4–5. Athleta (Neoathleta) noetlingi . Vredenburg 1923: 256; Cotter in Vredenburg 1928: 36 [new combination]. Lyria cossmanni (non Lyria cossmanni Vredenburg, 1923 ). Iqbal 1972: 65, pl. 19, figs 3, 9.

Comments. According to Cossmann & Pissarro (1909), the specimen illustrated on their pl. 2, figs 23–24 displays a biconic shape with spiny costae delineating a developed shoulder and looks more like a species of Volutospina than a species of Volutopupa Dall, 1890 (type species: Voluta cithara Lamarck, 1803 by original designation). We designate here this specimen as the lectotype of V. noetlingi . Vredenburg (1923) regarded Volutospina intercrenata Cossmann & Pissarro, 1909 as a junior synonym of V. noetlingi , considering that the specimens used to describe V. intercrenata represent juvenile specimens of V. noetlingi . However, the figured material of V. intercrenata does not display spiny costae delineating a developed shoulder, as observed in juvenile specimens of Volutospina species (e.g. the type species). Thus, these specimens do not belong to Volutospina and are not juveniles of V. noetlingi . In the Lakhra Dome, we have not found specimens of V. noetlingi , but a juvenile specimen of A. (V.) noetlingi from Jhirak has been illustrated by Iqbal (1972) under the erroneous name Lyria cossmanni . As in other juvenile specimens of Volutospina species, it has a biconic shape with spiny costae delineating a developed shoulder.

Stratigraphic range. Lakhra Formation: Jhirak ( Cossmann & Pissarro 1909; Vredenburg 1923).