Gigantotrichoderes cummingi Botero, Le Tirant, and Santos-Silva, 2018

Botero, Juan Pablo, Tirant, Stéphane Le & Santos-Silva, Antonio, 2018, A New Species of GigantotrichoderesTippmann, 1953 (Coleoptera: Cerambycidae: Cerambycinae: Torneutini) from Mexico, The Coleopterists Bulletin 72 (4), pp. 805-809 : 806-808

Gigantotrichoderes cummingi Botero, Le Tirant, and Santos-Silva , new species

( Figs. 3–7 ) Zoobank.org/ urn:lsid:zoobank.org:act:05C15C8C-2837-4EF8-9904-B5EBF1E3D0C6

Description. Male. Head dark brown except apex of antennal tubercles reddish brown, slightly lighter ventrally; palpi reddish brown; scape and pedicel dark brown; antennomere III dark brown basally, gradually dark reddish brown toward apex; remaining antennomeres reddish brown; pronotum and sides of prothorax dark brown except anterior and posterior margins of pronotum black; prosternum dark reddish brown, with prosternal process distinctly lighter toward apex; ventral side of meso- and metathoraces mostly dark reddish brown with dark brown areas interspersed; scutellum dark brown; elytra yellowish brown, gradually pale yellow toward apex, reddish brown on anterior inclined area, narrowly black along outer margin (gradually brown toward sutural angle); legs dark reddish brown with some areas darkened; abdominal ventrites dark reddish brown with some areas darkened. Head: Frons finely, densely punctate, slightly depressed on each side close to antennal tubercles; with yellowish bristly pubescence, moderately abundant, not obscuring integument, close to clypeus, dense on depressed areas, sparser and decumbent toward vertex. Area between antennal tubercles and upper eye lobes coarsely, shallowly, confluently punctate; with pale yellow pubescence not obscuring integument, denser laterally. Remaining surface of vertex coarsely, shallowly, confluently punctate along median groove, finely, densely rugose-punctate laterally; with pale yellow pubescence not obscuring integument. Median groove distinct from clypeus to between upper eye lobes, deeper between antennal tubercles and upper eye lobes; median groove continued as slightly elevated carina toward prothoracic margin. Area behind upper eye lobes finely, densely punctate close to eye, gradually slightly sparser toward prothorax; with pale yellow pubescence not obscuring integument near dorsal surface of head, gradually disappearing toward lower eye lobes. Area behind lower eye lobes smooth superiorly (this area narrowed toward ventral side), finely, abundantly punctate on narrow area close to eye toward ventral side, longitudinally striate on remaining surface; glabrous except narrow, pale yellow pubescent area close to inferior side of eye. Antennal tubercles finely, abundantly punctate basally, smooth on distal half; with moderately abundant, pale yellow pubescence basally, glabrous at apex. Genae short, triangular at apex, finely, sparsely punctate; with sparse, pale yellow pubescence interspersed with a few long erect setae of same color. Postclypeus finely, densely punctate on wide central area, smooth laterally; with short bristly, pale yellow pubescence on center of punctate area, distinctly longer, more abundant on sides of punctate area, glabrous on smooth area. Labrum small, slightly transversely depressed; finely, densely punctate; with long, erect, moderately abundant, reddish brown setae. Distance between upper eye lobes 0.46 times length of scape; in frontal view, distance between lower eye lobes 0.74 times length of scape; in ventral view, distance between lower eye lobes 0.75 times length of scape. Antennae slightly shorter than elytra, slightly surpassing base of posterior third of elytra, reaching about middle of abdominal ventrite III. Scape finely, abundantly punctate; outer surface convex; with pale yellow pubescence not obscuring integument. Dorsal surface of antennomeres with yellowish pubescence not obscuring integument. Antennal formula (ratio) based on length of antennomere III (not including projection of III–XI): scape = 1.75; pedicel = 0.40; IV = 1.13; V = 1.11; VI = 1.19; VII = 1.16; VIII = 1.13; IX = 1.19; X = 1.21; XI = 1.21; XII = 1.27. Thorax: Prothorax 1.3 times wider than long; divergent, slightly rounded from anterolateral angles to about middle, then subparallel-sided toward posterolateral angles; sides not tuberculate. Pronotum with 1 small, slightly elevated tubercle on each side of central area before midlength; moderately coarsely rugose-punctate, and minutely, densely punctate; with moderately dense, pale yellow pubescence, slightly less dense on centroposterior 2/3, especially on anterior region of this area. Sides of prothorax moderately coarsely rugose-punctate, with pale yellow pubescence not obscuring integument, sparser toward prosternum. Prosternum transversely tumid centrally; finely, slightly rugose-punctate, with sparse, pale yellow pubescence, denser close to procoxal cavities, central area, and close to anterior margin. Prosternal process slightly narrower than half of a procoxa, with apex slightly surpassing coxae; finely, abundantly punctate; with bristly, sparse, pale yellow setae, longer and more abundant laterally on central area. Ventral side of meso- and metathoraces with long, decumbent, abundant, pale yellow setae, denser on metathorax; mesoventral process about 0.75 times width of a mesocoxa, with sides of distal area slightly projectingd over coxae (tab-shaped). Scutellum with pale yellow pubescence not obscuring integument except glabrous posterior area. Elytra: Outer margins parallel-side; surface moderately coarsely, shallowly, abundantly punctate throughout; with minute, sparse, yellowish setae emerging from punctures, longer and denser along outer margins, especially toward apex; apex truncate, with outer angle rounded, and sutural angle with long spine. Legs: Femora sub-fusiform, metafemora more distinctly narrowed toward apex; with pale yellow pubescence not obscuring integument. Protibiae flattened dorso-ventrally, distinctly widened toward apex; meso- and metatibiae slightly flattened dorso-ventrally and less distinctly widened toward apex; all tibiae with yellowish pubescence not obscuring integument, longer, denser, bristly on inner side of meso- and metatibiae, especially on posterior half. Abdomen: Ventrites minutely, densely punctate except smooth posterior area on IV; with yellowish pubescence not obscuring integument, except glabrous smooth area on IV. Posterior margin of ventrite V wide, truncate, slightly concave centrally.

Dimensions. Holotype male: Total length = 30.60; prothoracic length = 5.25; posterior prothoracic width = 6.80; anterior prothoracic width = 4.50; maximum prothoracic width = 6.95; humeral width = 9.90; elytral length = 22.15.

Type Material. Holotype male from MEXICO, Veracruz: Sierra los Tuxtlas (900 m), 1–20.VII.2014, local collector ( MZSP).

Etymology. This species is dedicated to our friend and entomologist Royce T. Cumming.

Remarks. Gigantotrichoderes cummingi is similar to G. flabellicornis ( Fig. 8 ) but differs as follows: elytral reddish brown basal area not encroaching on dorsal surface; elytra without dark maculae on posterior half; prothorax somewhat truncated cone-shaped; central 2/3 of pronotum with distinct pubescence; pronotal punctures shallower. In G. flabellicornis , each elytron in both sexes has a distinct anterior, black macula encroaching on dorsal surface and projecting along sides in all specimens examined (n = 20); each elytron usually with distinct, dark maculae on posterior half; prothorax in males nearly parallel-sided on anterior 2/3; pronotal punctures deeper; and pronotum with large, glabrous or nearly glabrous, inverted Ushaped central area. The new species differs from G. conicicollis ( Fig. 2 ) by the shorter and stouter body (long and slender in males of G. conicicollis ), prothorax ( Fig. 3 ) without lateral tubercle (distinct lateral tubercle in males and female holotype of G. conicicollis ), and outer side of scape ( Fig. 6 ) convex (flattened in males of G. conicicollis ( Fig. 1 )). Gigantotrichoderes cummingi differs from G. brunneus by the elytra entirely light colored (entirely brown in G. brunneus ).

Figures

Figs. 1–8. Gigantotrichoderes conicicollis: 1) Scape and pedicel; 2) Male, dorsal habitus. Gigantotrichoderes cummingi, holotype male: 3) Dorsal habitus; 4) Ventral habitus; 5) Head, frontal view; 6) Scape and pedicel; 7) Lateral habitus. Gigantotrichoderes flabellicornis: 8) Male, dorsal habitus.