Coptocatus, Montandon, 1909

Dan, Polhemus, John, Polhemus & Sites, Robert, 2008, A Revision Of The Indochinese Genera Cheirochela And Gestroiella Heteroptera Naucoridae And A Review Of The Tribe Cheirochelini, Raffles Bulletin of Zoology 56 (2), pp. 255-279 : 257

COPTOCATUS Montandon, 1909

Diagnosis. – Form elongate-oval. ( Fig. 1 ) Abdominal and thoracic venter with fine pile of short, closely appressed hairs. Pronotum not markedly expanded posteriorly (except in C. stereos ), posterior width less than 2× width of head; anterolateral angles of pronotum simple, not forming ventrallydirected cup-shaped depressions at apices. Hemelytra of brachypterous forms short and truncate, apices rounded, not reaching to abdominal tergite VI. Parameres variable in shape, either deeply cleft medially and asymmetrically bidentate, or uniramous and tapering on distal half, sometimes with a fringe of long setae on outer margins.

Discussion. – The genus Coptocatus is confined to Borneo, and was proposed by Montandon (1909) to hold a single species, C. oblongulus , collected near Mt. Kinabalu. Five additional species have been described subsequently by Polhemus (1986), Nieser & Chen (1991) and Zettel (2002). Although clearly members of the tribe Cheirochelini based on synapomorphies of the head and ventral abdomen (see Polhemus, 1986, Figs. 7, 8 ), Coptocatus species are narrower and more elongate in comparison to those of Gestroiella or Cheirochela (compare Figs. 1 , 2 and 18 ). The key below, modified from Zettel (2002), will permit identification of the species described to date.

Figures

Fig. 1. Coptocatus kinabalu Polhemus. Male, dorsal habitus. Drawn by Young Sohn.

Figs. 7–11. Cheirochela species, male right paramere: 7, C. assamensis, holotype; 8, C. feana, paratype. 9. C. birmaniensis, holotype; 10, C. tonkina, paratype; 11, C. thailandana, paratype.

Fig. 2. Cheirochela tonkina Polhemus & Polhemus. Male, dorsal habitus. Drawn by Young Sohn.

Fig. 18. Gestroiella siamensis new species, female, dorsal habitus. Drawn by Young Sohn.