Odontanax Lee, Satô, and Yang, 2000

Shepard, William D. & Sites, Robert W., 2019, Larval Psephenidae (Coleoptera: Byrrhoidea) of Thailand: Annotated List and Illustrated Key to Genera, The Coleopterists Bulletin 73 (2), pp. 259-282 : 264-265

Odontanax Lee, Satô, and Yang, 2000 Fig. 29 , Map B

Discussion. Odontanax contains 12 species that are distributed from Sri Lanka, India, and Nepal east to China and south to Malaysia, Indonesia, and the

Psephenidae Lacordaire, 1854

Eubrianacinae Böving and Craighead, 1931 Jaechanax Lee, Satô, and Yang, 2000 major (Pic, 1913) Jinbrianax Lee, Satô, and Yang, 1999 metallicus ( Pic, 1922) schillhammeri Lee, Satô, and Yang, 1999 Mubrianax Lee, Satô, and Yang, 1999 * Odontanax Lee, Satô, and Yang, 2000 maculicollis (Fairmaire, 1888) thai Lee, Satô, and Yang, 2000 undescribed larva *

Eubriinae LeConte, 1861 Dicranopselaphus Guérin-Méneville, 1861 bicolor Lee and Yang, 1996 doiinthanonus Lee and Yang, 1996 fangensis Lee and Yang, 1996 rufus (Pic, 1916) yamasakii Lee and Yang, 1996 Granuleubria Jäch and Lee, 1994 satoi Lee and Yang, 1999 Homoeogenus Waterhouse, 1880 * Macroeubria Pic, 1916 Bella Lee, Yang, and Satô, 1999 contracta Lee, Yang and Satô 1999 diffusa Lee, Yang, and Satô, 1999 flabellata Lee, Yang, and Satô, 1999 fulva Lee, Yang, and Satô, 1999 impressicollis Pic, 1934 Microeubria Lee and Yang, 1994 hajeki Lee, 2016 longicornis Lee and Jäch, 1996 Schinostethus Waterhouse, 1880 flabellatus Lee, Yang, and Brown, 1993 malickyi Lee, Yang, and Satô, 1998 nigricornis Waterhouse, 1880 notatithorax (Pic, 1923)

Psepheninae LeConte and Horn, 1883 Mataeopsephus Waterhouse, 1876 *

Psephenoidinae Bollow, 1938 Afropsephenoides Basilewsky, 1959 Heteropsephenoides Jeng and Jäch, 2003 horaki Jeng and Jäch, 2003 Micreubrianax Pic, 1954 siamensis Jeng, Jäch, and Yang, 2006 Psephenoides Gahan, 1914 Sinopsephenoides Yang, 1993 malickyi Jäch and Jeng, 1995

Philippine Islands. Adults are often collected at lights. In Thailand, Odontanax maculicollis (Fairmaire, 1888) , and Odontanax thai Lee, Satô, and Yang, 2000 are reported from Mae Hong Son Province: Ban Si Lang ( Lee et al. 2000a), Chiang Mai Province: Fang, and Mae Hong Son Province: Ban Si Lang ( Lee et al. 2000a).

Material Examined. Chanthaburi Province: Khao Soi Dao, Wildlife Sanctuary; stream, from Namtok Khao Soi Dao , 13°5.810’N 102°10.332’E, 523 m, 17 January 2016, L-1911, R. W. Sites (1L) . Chiang Mai Province: creek at Chiang Dao, Wildlife Research Center, 19°21’N 98°55’E; 520 m, 26 Aug 2002, coll: CMU team (3L); Doi Inthanon National Park , Mae Pan River @ Mae Chaem dist. , 540 m, 18°31’N 98°23’E, 23 IX 02, P. Thamsenanupap (4L); Doi Inthanon National Park , Mae Klang River at Namtok Sob Ab; 18°31’N 98°36’E, 543 m, 17 May 2004, L-691, Laudee & Thamasenanupap (1L); Doi Inthanon National Park , Siriphum Waterfall, 1380 m, 18°32’N 98°31’E, 2 III 2002, D. Tapanva (1L); same locality except level 2, 1460 m, 18°32’N 98°31’E, 21 III 2002, L-314, R. W. Sites (1L); Doi Sutep-Pui National Park , creek from Mohk Fah waterfall, 564 m, 19°06’N 98°46’E, 18 March 2002, L-304, Sites, Vitheepradit, Kirawanich (2L) . Chiang Rai Province: stream from Khun Kon waterfall, 524 m, 19°51’N 99°39’E, 19 April 2003, L-466, AV, Prommi, Setaphan (2L) . Kanchanaburi Province: Ampur Sai Yok, Thong Pha Phum Reforestation Station, Mae Nam Noi , 204 m, 14°31’N 98°37’E, 12 April 2002, L-335, UMC and CMU teams (1L); Amphur Sangkhla Buri, Huay Kob, 289 m, 15°13’N 98°22’E, 13 IV 2002, L-339, UMC and CMU teams (2L); Huay Ka Yaeng at Ban Pa Soo Klang, rocky stream, 751 m, 16°22.370’N 101°22.777’E, 31 May 2011, L-1297, R. W. Sites (4 L); Amphur Thong Pha Phum, Huay Ou Long, 139 m, 14°46’N 98°40’E, 11 Apr 2002, L-332, UMC and CMU teams (1L); Huay Ou Long at Ban Ou Long, gravel stream, 274 m, 14°35.177’N 98°34.921’E, 1 June 2011, L-1301, R. W. Sites (3L); Huay Ou Long, opposite Keong Ka Wia Reforestation Stn., gravel & vegetative margins, 326 m, 14°33.199’N 98°34.238’E, 19 January 2015, L-1872, R. W. Sites, A. Vitheepradit (1L); same locality, 1 June 2011, L-1300, R. W. Sites (1L); Thong Pha Phum National Park , Huay Pak Kog, 139 m, 14°47.183’N 98°48.401’E, 1 June 2011, L-1303, R. W. Sites (3L) . Khampaeng Phet Province: Khlong Lan Natl. Park, Namtok Khlong Lan, 310 m, 16°07’N 99°16’E, 12 Aug 2002, CMU team (4L); same locality, 7 April 2003, L-453, Sites, Vitheepradit, Prommi and Setaphan (1L); same locality, 20 VIII 2002, coll. Arthit (1L); same data except 26 IX 2002, coll. Arthit (1L); same data except 23 XII 2002, coll. Arthit (1L); same data except 30 V 2003, coll. Arthit (3L); Khlong Lan Natl. Park, Khlong Nam Lai Waterfall, 16°11’N 99°15’E, 751 m, 6 April 2003, L-448, Sites, AV, Prommi, Setaphan (4L); Khlong Lan National Park , Phetcha Khor Waterfall, 270 m, 16°17’N 99°14’E, 7 April 2003, L-452, Sites, AV, Prommi, Setaphan (1L) . Nakon Nayok Province: Khao Yai National Park, ck. @ ‘km 29’, 14°22’N 101°25’E, 11 VIII 1997, G. W. Courtney (2L); Khao Yai National Park, Salika Waterfall , gravel, 68 m, 14°18’N 101°15’E, 7 April 2004, L-606, Sites & Vitheepradit (1L); Wang Takrai WS, 43 m, 14°19.689’N 101°18.262’E, 8-9 iii 2012, R. W. Sites (1L) . Nakon Ratchasima Province: Khao Yai National Park , 200 m E of Heaw Narok Waterfall, 397 m, 14°17’N 101°23’E, 5 April 2004, L-602, Sites & Vitheepradit (1L) . Nakhon Si Thammarat Province: Nopphitam, Ban Pitam, 6.7 km SW int. 4186 X 4188, Tumbon Krung Ching , gravel strm, 98 m, 08°44’N 99°39’E, 30 April 2002, L-382, Vitheepradit, Kirawanich (1L); Nopphitam Khlong Yod Leung, 78 m, 26 May 2005, L-800, Sites, Vitheepradit & Prommi (1L); Tumbon Krung Ching, Klong Nob, 89 m, 08°44’N 99°39’E, 25 May 2005, L-798, Sites, Vitheepradit, Prommi (2L) . Nan Province: Amphur Bo Kluea, Tumbon Phuka , Doi Phuka Park Ranger Station 6, Huey Korg , stream, gravel, 516 m, 19°01’N 101°11’E, 22 April 2003, L-474, AV, Prommi, Setaphan (1L); Mae Charim National Park , Nam Wa River , 335 m, 18°36’N 100°59’E, 23 April 2003, L-478, AV, Prommi, Setaphan (2 L) . Narathiwat Province: stream below Bacho Waterfall , 62 ft, 6°29’ 56.28”N 101°38’ 10.16”E, 15 Jan 1995, L-78, Sites, Nichols & Permkam (1L) . Pattani Province: Sai Khao , 1015 ft, 6°39’06.21”N 101°05’ 56.86”E, 16 Jan 1995, L- 80, Sites & Nichols (3L) . Petchabun Province: 5 km E Sila, Mae Nam Pa Sak, 2 III 1994 (WDS- A-1025 on reverse), William D. Shepard, leg. (2L); same locality, 3 IV 1994, (WDS-A-1050 on reverse), William D. Shepard, leg. (3L); Nam Nao Nat. Park , 22 III 1994, Prom Laeng (WDS-A- 1040 on reverse), William D. Shepard, leg. (2L) . Petchaburi Province: Amphur Tha Yang, Ban Yang Chum , stream, 46 m, 12°47’N 99°40’E, 15 May 2003, L-536, Vitheepradit & Ferro (2L); Huay Sat Yai, sandy stream near ocean, 109 m, 12°30.824’N 99°34.152’E, L-1999, RWS (2L); Kaeng Ka Chan N. P., Pranburi stream at Ban Krang, 380 m, 12°48.037’N 99°26.557’E, 28 May 2004, L-721, Vitheepradit & Prommi (50L) . Phang Nga Province: Sri Phang Nga National Park, Tam Nang Waterfall , stream, gravel, 68 m, 8°58’N 98°27’E, 8 June 2006, L-926, Sites, Vitheepradit, Prommi (5L) . Phayao Province: Doi Luang National Park, Namtok Cham Pa Thong, 620 m, 19°13’N 99°44’E, 27 Mar 2003, L-420, Sites, Vitheepradit, Prommi (1L); Huay Chom Poo Wildlife Sanctuary Station , 526 m, 19°10’N 100°06’E, L-468, Vitheepradit, Prommi, Setaphan (1L) . Phrae Province: Wieng Ko Sai National Park, Namtok Maekueng, tier 1, 350 m, 17°58’N 99°35’E, CMU team (9L); Wieng Ko Sai National Park , upper Nam. Punjane , 295 m, 17°56’N 99°34’E, 20 Feb 03, CMU team (1L); Wieng Ko Sai National Park , Namtok Punjane (lower), 295 m, 17°57’N 99°34’E, 22 VII 2002, CMU team (1L); same locality, 285 m, 17 May 2003, CMU team (3L) . Prachin Buri Province: Khao Yai National Park, Ta Krow Waterfall , margin, 26 m, 14°11’N 101°35’E, 8 April 2004, L-609, Sites & Vitheepradit (4L) . Prachuap Khiri Khan Province: Amphur Hua Hin, Tumbon Huay Pheung , Ban Sak Yai , 121 m, 12°30’N 99°34’E, 21 Apr 2002, L-358, Vitheepradit & Kirawanich (1L); same locality except, stream, 114 m, 12°29’N 99°34’E, 16 May 2002, L-537, Vitheepradit & Ferro (4L); Amphur Hua Hin, 10.5 km E of Nam Tok Pa La Or on Hwy 3219, 134 m, 12°31’N 99°33’E, 20 April 2002, L-356, Vitheepradit & Kirawanich (1L); Amphur Kui Buri Forest Plantation Station , gravel and rock stream, 117 m, 12°04’N 99°37’E, 29 May 2002, L-725, Vitheepradit & Prommi (1L); same data except 21 May 2005, L-785a, Sites, Vitheepradit & Prommi (7L); Kaeng Krachan N. P., Pa La Ou Waterfall , gravel, 319 m, 12°32’N 99°27’E, 28 May 2004, L-722, Vitheepradit & Prommi (5L) . Ratchaburi Province: Amphur Suan Phueng , 14.6 km SSE Suan Phueng, stream, 183 m, 13°28’N 99°15’E, 17 IV 2002, Vitheepradit, Kirawanich (1L) . Sa Kaeo Province: Pang Sida Natl. Park, Klong Huay Nam Yen , 218 m, 14°02’N 102°15.955’E, 16 April 2004, L-630, Sites & Vitheepradit (2L) . Sakon Nakhon Province: Phu Pan N. P., Huay Teuy , 253 m, 17°04.770’N 103°57.835’E, 26 April 2004, L-645, A. Vitheepradit (1L) . Songkhla Province: Ton Nga Chang Natl. Park, stream at Buddhist temple, 44 m, 6°56’N 100°15’E, 6 July 1997, L-127, R. W. Sites (4L) ; Ton Nga Chang Wildlife Sanct., Ton Plieuw Waterfall, 120 m, 7°00’N 100°14’E, 17 June 2006, L-957, Vitheepradit & Prommi (1L); Khao Nam Khang National Park, 102 km SE of Amphur Hat Yai, Amphur Na Tawee, 100 m, 6°36’N 100°36’E, L-246, R. W. Sites (1L); Khao Nam Chang , ca. 21 km SW Nathawai, 6°34’N 100°40’E, 13 January 1995, L-72, Sites & Nichols (2L) . Suratthani Province: Amphur Phanom, Ban Chong Lom , Klong Sok , 43 m, 08°52’N 98°40’E, 23 May 2003, L-563, Vitheepradit & Ferro (1L); Tai Rom Yen National Park , Dard Fah Waterfall , gravel, 236 m, 08°51’N 99°28’E, 17 June 2004, Sites, Vitheepradit & Prommi (1L) . Uttaradit Province: Mae Cheuy Waterfall , 231 m, 17°47’N 100°10’E, 27 April 2003, L-487, AV, Prommi, Setaphan (1L) . Yala Province: Than To, Banglang National Park , 6°09’N 101°11’E, 14 Jan 1995, L-73, Sites & Nichols (1L) .

Figures

Figs. 25–30. Eubrianacinae genera, dorsal habitus. 25) Eubrianax; 26) Jaechanax; 27) Jinbrianx; 28) Mubrianax;