Ampullaria chiquitensis, d'Orbigny, 1838

Cowie, Robert H. & Héros, Virginie, 2012, Annotated catalogue of the types of Ampullariidae (Mollusca, Gastropoda) in the Muséum national d’Histoire naturelle, Paris, with lectotype designations, Zoosystema 34 (4), pp. 793-824 : 800

chiquitensis d’Orbigny, 1838 , Ampullaria

Ampullaria chiquitensis d’Orbigny, 1838 (6 May): pl. 48, figs 10, 11 [1838 (11 June): 367].

CURRENT TAXONOMIC STATUS. — Marisa . Synonym of Helix cornuarietis Linnaeus, 1758 (see Remarks).

TYPE SPECIMEN. — Holotype, the shell illustrated by d’Orbigny (1838: pl. 48, figs 10, 11) was not found in MNHN.

TYPE LOCALITY. — “sud-est de la province de Chiquitos (république de Bolivia) […] entre les Missions de San- Miguel et de San-José […] à peu près de la première Mission, dans le lac de los Migueleños, et dans les marais des environs”.

REMARKS

Although we have found no type material in the MNHN, we include this entry in our catalog in order to correct the interpretation of Cowie & Thiengo (2003: 50) regarding other material in both the MNHN and NHMUK, formerly considered as syntypes. Plate 48 was published on 6 May 1838 and page 367 on 11 June 1838 ( Cowie & Thiengo 2003: 91). The plate (with name) therefore confers availability of the name from the former date. The two illustrations appear to be of the same shell, which is therefore the holotype, by monotypy. Other shells mentioned in the subsequent text are irrelevant and have no standing in nomenclature, as they were not in the publication conferring availability of the name (the plate with name). The MNHN 4479 shell, previously treated as a syntype, is not the shell illustrated; based on the banding pattern, neither is any of those in the NHMUK. Therefore, contrary to Cowie & Thiengo (2003: 50), none of these specimens has any type status. Cowie & Thiengo (2003: 50), based on previous works, treated this taxon as a synonym of Helix cornuarietis . However, based on the distributions of M. cornuarietis and M. planogyra Pilsbry, 1933 (Hayes 2009; Hayes et al. 2009a, b), and the type locality of Ampullaria chiquitensis , it is more likely that A. chiquitensis is a synonym of M. planogyra , although this issue requires further research and we refrain from formally synonymising them here.