Ampullaria baeri, Dautzenberg, 1902

Cowie, Robert H. & Héros, Virginie, 2012, Annotated catalogue of the types of Ampullariidae (Mollusca, Gastropoda) in the Muséum national d’Histoire naturelle, Paris, with lectotype designations, Zoosystema 34 (4), pp. 793-824 : 796

baeri Dautzenberg, 1902 , Ampullaria

( Fig. 1B )

Ampullaria baeri Dautzenberg, 1902: 312 , pl. 9, figs 12, 13.

CURRENT TAXONOMIC STATUS. — Pomacea (Effusa) . Valid species.

TYPE SPECIMEN. — Lectotype (here designated) MNHN 4493 .

TYPE LOCALITY. — “ Rio Mixiollo [= Misciotto; Berthold 1991: 13], province de Huallaga, Pérou ” .

REMARKS

Fischer-Piette (1950) indicated a holotype (“ 34 mm ”), which Cowie & Thiengo (2003) and Köhler & Glaubrecht (2006) considered to have constituted an inadvertent lectotype designation. Dautzenberg implied that his description was based on more than one specimen by his use of “parfois” (“sometimes”) in reference to certain shell characters, which we interpret as meaning in some specimens. The ICZN (1999b; art. 74.5) says that: “When the original work reveals that the taxon had been based on more than one specimen [which this description does], a subsequent use of the term ‘holotype’ [as by Fischer-Piette (1950)] does not constitute a valid lectotype designation unless the author [i.e. Fischer-Piette (1950)], when wrongly using that term, explicitly indicated that he or she was selecting from the type series that particular specimen to serve as the namebearing type.” Fischer-Piette (1950) made no such explicit indication, so his simple listing of the single specimen as the “ holotype ” was not a valid lectotype designation, contrary to the interpretation of Cowie & Thiengo (2003) and Köhler & Glaubrecht (2006). The shell is 34 mm maximum width, agreeing with Fischer-Piette (1950). The illustration is a little smaller and does not quite match in shape, but the labels, including one in Dautzenberg’s hand, say that it is the illustrated shell. We here designate it as the lectotype to stabilise the name.

In shape, this shell (although smaller) corresponds closely to many specimens of Helix glauca Linnaeus, 1758 seen by the first author, as well as to a specimen of Nerita effusa Müller, 1774 , labeled in the MNHN collections as from the Lamarck collection, and to the probable syntype of Ampullaria gevesensis Deshayes, 1838 in the MNHN type collection, both of which are synonyms of Helix glauca Linnaeus, 1758 , according to Cowie & Thiengo (2003). However, this shell (which retains its periostracum) is fairly uniformly tan in colour and has an orange peristome, features seen consistently in other non-type material and that do not appear to intergrade with other shells in the MNHN, nor with other material conventionally referred to Helix glauca Linnaeus, 1758 . Although Boss & Parodiz (1977: 116) suggested that it is probably synonymous with glauca Linnaeus, 1758 , the two taxa have never been formally synonymised and are probably distinct.

Figures

FIG. 1. — A, Ampullaria aulanieri Deville & Huppé,1850,lectotype MNHN 23072, 48.2 mm;B, Ampullaria baeri Dautzenberg,1902,lectotype MNHN 4493, 27.0 mm; C, Ampullaria begini Morlet, 1889,syntype MNHN 4499, 21.4 mm; D, Ampullaria belizensis Crosse & Fischer in Fischer & Crosse, 1888, holotype MNHN 5139, 61.8 mm; E, Meladomus bloyeti Bourguignat, 1889, lectotype MNHN 4501, 63.3 mm; F, Ampullaria bruguieri Deshayes, 1830, lectotype MNHN 23074, 72.1 mm; G, Ampullaria carinata Olivier, 1804, lectotype MNHN 23075, 28.5 mm; H, Ampullaria castanea Deshayes, 1830, lectotype MNHN 4710, 42.2 mm.