Parenypnias inauditus, Schwarzhans & Klots & Ryabokon & Kovalchuk, 2022

Schwarzhans, Werner, Klots, Oleksandr, Ryabokon, Tamara & Kovalchuk, Oleksandr, 2022, A rare window into a back-reef fish community from the middle Miocene (late Badenian) Medobory Hills barrier reef in western Ukraine, reconstructed mostly by means of otoliths, Swiss Journal of Palaeontology (18) 141 (1), pp. 1-35 : 13

publication ID	https://doi.org/ 10.1186/s13358-022-00261-3
DOI	https://doi.org/10.5281/zenodo.13127108
persistent identifier	https://treatment.plazi.org/id/039D7D5B-FE45-FFAE-FBC8-FEB0FC7AFCD2
treatment provided by	Felipe
scientific name	Parenypnias inauditus
status	sp. nov.

Treatment

Parenypnias inauditus n. sp.

Figure 5e–h View Fig

2020a Ponticola sp. —Schwarzhans, Brzobohatý & Radwańska: pl. 4, fig. 18.

Holotype NMNHU-P PI 2550 , Kozatskyi Yar , western Ukraine, late Badenian, Fig. 5f. View Fig

Paratypes 4 otoliths: 3 specimens, Shydlivshchyna , NMB P1207 View Materials ; 1 specimen, late Badenian , Žižkov 1 well (1616– 1622 m), Czech Republic, DGS MU-0395 .

Etymology From inauditus (Latin) = unbeknown, referring to the occurrence of the species outside of the extant distribution range of the Gobiosomatini.

Diagnosis OL:OH = 1.15–1.22. Preventral projection pointed; postdorsal projection moderately long, pointed, slightly to moderately bent outward. Dorsal rim regularly curved but with pointed middorsal denticle. Sulcus narrow, short (OL:SuL = 2.2–2.5), inclined at 12–15°. Ostial lobe absent or very weak; no subcaudal iugum. Inner face equal or less bent than outer face.

Description Moderately slender, moderately thin, small otoliths reaching 1.6 mm in length (holotype). OH:OT = 2.7–3.0. Anterior rim inclined at 77–83°, with distinct concavity at about level of ostial tip and distinct, mostly sharp and moderately long preventral projection. Predorsal angle distinct, angular, somewhat projecting, positioned moderately high on anterior rim. Ventral rim nearly straight, smooth, slightly undulating in smallest specimen ( Fig. 5h View Fig ). Dorsal rim overall relatively regularly curved, sometimes slightly undulating ( Fig. 5h View Fig ), with distinct angular or projecting denticle in middorsal position, with distinct moderately strong, sharp postdorsal projection, slightly to moderately bent outward. Posterior rim with distinct incision below postdorsal projection and variably developed postventral angle.

Inner face slightly convex to nearly flat, with very narrow, slightly deepened, centrally positioned sulcus. SuL:SuH = 2.6–3.2. Sulcus without discernable ostial lobe, oval-shaped, with rounded tips, inclined at 12–15°. No subcaudal iugum. Ventral furrow broad, distinct, reaching from anterior to posterior tips of sulcus, curvature more or less following ventral rim of otolith. Dorsal field with small, indistinct dorsal depression, ventrally marked by feeble crista superior toward sulcus and dorsally closed to dorsal rim of otolith. Outer face convex, equally or more convex than inner face, regularly shaped, smooth.

Discussion Parenypnias inauditus is easily recognized by certain features of the outline of the otolith, such as the sharply pointed preventral projection, the pointed postdorsal projection, and the middorsal denticle, which, however, is easily affected by erosion and also seems to be developed more like a middorsal angle in specimens smaller than 1.25 mm in length ( Fig. 5g, h View Fig ). It differs from the parallel occurring P. kiselevi n. sp. described below in the longer sulcus (OL:SuL = 2.0–2.5 vs. 2.0), the more pronounced preventral projection, the less strongly outward bent postdorsal projection, the sharp otolith rims (vs. thick), and the more slender shape (OL:OH = 1.15– 1.22 vs. 1.05).