MENGYANGENSIS, Daugeron & Grootaert, 2005

Daugeron, Christophe & Grootaert, Patrick, 2005, Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae), Zoological Journal of the Linnean Society 145 (3), pp. 339-391 : 377-378

EMPIS (COPTOPHLEBIA) MENGYANGENSIS SP. NOV.

( FIG. 32 )

Type material: Holotype: male, China, Yunnan Province, Jinhong, Mengyang , 11.iii.1999, primary rainforest, P.Grootaert (sample 99090) ( KBIN –IRScNB) . Paratypes: 3 males, same data but 5.iii.1999 (sample 99060) ( KBIN –IRScNB) ; 3 males, 2 females, same data than holotype (sample 99091) ( KBIN –IRScNB) .

Other material: Because of their poor condition the following specimen is not considered paratype: 1 male, same data than holotype but 5.iii.1999 (sample 99060) ( KBIN –IRScNB) .

Etymology: The name of the species is derived from the type-locality.

Description: Blackish to yellowish species of middle size, yellowish legs, male fore tibia deformed with several anterodorsal spines, hind tarsus very thick, black with long bristles, phallus with small ventral notch at middle. MALE: HEAD. Occiput dusty brown-grey. Face blackish, dusty. Palpus brownish. Antenna black, first flagellomere conical, second and third flagellomeres aristiform. Labrum brownish, length about twice head height. Labium blackish, with short bristles. THORAX dark brown in ground colour. Prescutellar depression strongly dusty. All bristles black. Antepronotum with short lateral bristles. Proepisternum and prosternum apparently with 1 and 2 bristles, respectively. Acrostichals irregularly biserial, fine, short. Dorsocentrals apparently uniserial, ending with 2 strong, long bristles in prescutellar depression. Anterior and posterior spiracles dark brown. LEGS ( Fig. 32B ). Coxae, femora yellow, tibiae yellowish to brownish, tarsomeres blackish. Fore tibia deformed, strongly narrowed at base, with 6 spine-like, short dorsal bristles on basal third, 1 rather strong, long dorsal bristle almost at middle; first fore tarsomere somewhat swollen, with pair of dorsal bristles apically. Mid femur with long ventral bristles on apical half; mid tibia with 3 strong, long dorsals, 1 strong, long ventral at basal third; first mid tarsomere with 1 strong, long dorsal apically. Hind femur somewhat S-shaped in dorsal view, with distinct posterolateral bristles basally; hind tibia distinctly dilated apically, interiorly curved, with row of strong, long dorsals; hind tarsomeres swollen, with strong, long dorsals; first hind tarsomere with two ventral rows of short, strong bristles. WING (length = 2.75) clear. Halter with yellow base, dark brown knob. ABDOMEN with first four tergites brown, first five sternites yellow, otherwise dark brownblackish. Distinct bristles at base, posterior margin of sternites with pair of strong, long bristles, also on tergites posteriorly and laterally. Segment 8 with strong, long posterior bristles. Hypopygium ( Fig. 32A ). Cercus with minute spines at tip. Epandrium somewhat rounded at tip, with pair of small anterodorsal protuberances bearing several long bristles; c. 12 strong, long bristles ventrally and apically. Hypandrium well sclerotized, sickle-shaped in lateral view. Phallus slender, with small ventral notch at middle.

FEMALE: Similar to male except for following characters: lower ommatidia enlarged, frons slightly wider than face, brownish-dusty grey. Legs with shorter bristles, mid femur with dorsal pennation except apically, shorter ventral pennation at apical half; mid tibia with flattened dorsal bristles apically; hind femur with long dorsal and ventral pennation; hind tibia with long dorsal pennation, long ventral pennation at apical half; first hind tarsomere with dorsal pennation except apically. Abdomen distinctly brown-yellowish, darker on last three segments.

Figures

Figure 32. The Empis (Coptophlebia) jacobsoni clade, E. (C.) mengyangensis sp. nov. A, male hypopygium in lateral view. Scale bar = 0.1 mm. B, male hind leg (tibia and tarsus) in lateral view. Scale bar = 0.5 mm.