Limonium carpathum

Brullo, Salvatore & Erben, Matthias, 2016, The genus Limonium (Plumbaginaceae) in Greece, Phytotaxa 240 (1), pp. 448-450 : 448-450

20. Limonium carpathum (Rech. fil.) Rechinger fil. (1943a: 427) ( Fig. 20 )

Basionym :― Statice carpatha Rechinger fil. (1936: 147). Type (lectotype, designated here):― GREECE. Dodecanese : Insula Karpathos (Scarpanto): In saxosis calc. litoreis ad Vrondi adversus Pigadia, 17 June 1935, K. H. & F. Rechinger 8241 (lectotype G!, isotypes B!, M!, W!, WU!) .

Description: —Plant perennial, glabrous, forming a sub-shrub 30–40 cm tall, with several ascending to erect stems and a robust tap-root. Caudices 2–12 cm long, branched, densely spirally leafy in the upper half. Leaves thick, coriaceous, grey-glaucous, 50–140 mm long and 1.0– 3.5 mm broad, linear-oblanceolate, apex acute to obtuse, verrucate to tuberculate with crateriform glands, with one central nerve, gradually tapering into the petiole. Stems slender, 10 –50 cm long, verrucate to tuberculate with crateriform glands, nearly straight, grey-glaucous, slightly flexuous, branching frequently begins upper the lower third. Inflorescence narrowly trullate in outline. Sterile branches absent or 1–3 per stem, 1.5–3.0 cm long, unbranched. Fertile branches 3–10 cm long, flexuous, directed obliquely upwards, forming branching angles of 20°–45°, normally unbranched. Spikes 30–150 mm long, straight to slightly flexuous, erect to directed obliquely upwards. Spikelets 7.4–8.2 mm long, composed of 1–2 (–3) flowers, very remotely arranged with 1–4 per 2 cm. Outer bract 1.9–2.5 mm long and 1.9–2.3 mm broad, triangular-ovate, obtuse; bract margin broadly membranous; central part slightly fleshy, sometimes only at the base, acuminate. Middle bract membranous, 2.1–2.6 mm long and 1.6–2.0 mm broad, oblong-elliptic, rounded. Inner bract 5.8–7.0 mm long and 3.2–4.0 mm broad, obovate-elliptic, rounded; bract margin broadly membranous; central part fleshy, 4.8–5.2 mm long and 2.1–2.6 mm broad, oblong to oblong-obovate, acuminate, forming a narrowly triangular tip, 1.0– 1.2 mm long, not reaching the upper margin. Calyx 5.0–6.0 mm long, exceeding the inner bract by 0.5–1.0 mm; calyx tube glabrous or very sparsely shortly hairy, with 5 ribs ending near the upper margin of lobes; calyx lobes fine, ca. 0.4 × 0.9 mm, triangular. Corolla lilac.

Chromosome number: —2n = 34 (Erben Li–1767, unpubl.)

Distribution: —Restricted to Karpathos, where it is very frequent ( Fig. 92B ).

Habitat: —Calcareous and marly rocky coast.

Taxonomic remarks: — L. carpathum is very similar to L. sitiacum and L. monolithicum , but it differs from both by having nearly needle-like leaves and triangular calyx lobes.

Specimens examined: — GREECE. Dodecanese: Insula Karpathos (Scarpanto), in saxosis calc. litoreis ad Vrondi adversus Pigadia, 17 June 1935, K. H. & F. Rechinger 8241 (BM, M); Insula Karpathos (Scarpanto), in saxosis calc. litoreis ad Phiniki (Finiki), 18 June 1935, K. H. & F. Rechinger 8323 (BM, G, W); Insula Karpathos, ad portum vici Vrondi, alt. 5– 15 m. In rupestribus calcareis ad litus, 5 June 1963, Greuter 5822 (M, W, Herb. Greuter); Karpathos (Griechenland), E-Küste am Kap Patella, 3 km südöstl. Pigadhia, Steilküste im Salzwasser-Spritzbereich, 21 April 1982, Raus 5667 & Pleger (B); Karpathos (Griechenland), Tal von Ahata (östli. Aperi), steile Küstenfelsen, Kreidekalk, 0–40 m, 19 May 1982, Greuter 19196, Pleger & Raus 6508 (B, Herb. Greuter); Insula Karpathos, Pigadia, ad litus pone vicum Vrondi, secus centrum vis electricae, alt. 5–10 m, in rupestribus calcareis litoreis, 25 August 1982, Greuter & B. Zimmer 19545 (B, C, G, MSB 88121); Karpathos, 2 km nordwestl. von Ammopi, flache Felsküste, 5–10 m, 14 May 1983, Raus 8226 (B, C); Karpathos Lefkos (bei der Insel Sokastro), Sandstrand und flache Felsküste, 2–10 m, 24 May 1983, Raus 8392 (B); Karpathos, Ostspitze der Insel, Kalkgestein, 5 m, 27 May 1983, Raus 8450 (B); Karpathos, 0.5 km südöstl. von Lefkos, flache Felsküste mit anschliessender Kleinstrauchvegetation, 10 m, 25 May 1983, Raus 8470 (B); Karpathos, Ausgang des Tales von Kira Panagia östlich Aperi, Küstenfelsen und Sandstrand, 2–10 m, 16 June 1983, Raus 8806 (B); Karpathos, Westküste gegenüber der Kleininsel Proni, Felsstrand, Kalkgestein, 5–15 m, 22 May 1984, Raus 9905 (B, C); Karpathos, W-Küste der Insel, nahe der Kleininsel “Proni”, ausgeprägte Halophytenflur und Phrygana, 0–40 m, 14 April 1986, Hoener & Potthoff 271 (B); Karpathos, Küstenfelsen südlich Mesochori, 5 m, 26 May 1994, Ulrich s.n. (M); Karpathos, Höhen über der Bucht zwischen Kap Volakas und Amopi, 0–40 m, Strandfelsen, 6 April 1995, Kalheber 95–073 (Herb. Kalheber); Karpathos, O-Seite, Achata-Bucht, Sandstrand und Felspartien, 0–30 m, 10 April 1995, Kalheber 95–465

LIMONIUM (PLUMBAGINACEAE)

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BRULLO & ERBEN

(Herb. Kalheber); Karpathos, Kira Panagia, Strandbereich der Bucht, 0–15 m, 12 April 1995, Kalheber 95–586 (Herb. Kalheber); Insel Karpathos, Ahata Strand, N Siete, Felsen in Küstennähe, Kalk, 5 m, 26 April 1995, Jahn s.n. (B); Karpathos, Pigadia, 28 August 1996, Brullo & Guarino s.n. (CAT, Herb. Erben); Karpathos, Poseidon, 28 August 1996, Brullo & Guarino s.n. (CAT); Isola di Karpathos, Achata, 29 August 1996, Brullo & Guarino s.n. (CAT, Herb. Erben); Karpathos, Finiki, 30 August 1996, Brullo & Guarino s.n..sheet 1 (CAT, Herb. Erben); Karpathos, Felsplateau zwischen dem Ormos Amorfos und dem Ort Amopi, 15–30 m, Felsphrygana, 1 June 1998, Kalheber 98–785 (Herb. Kalheber); Karpathos, Ostküste, zwischen Kathodio und Kira Panagia, Felsen am Strand in Kira Panagia, 11 June 1998, Kalheber 98–1183 (Herb. Kalheber); Karpathos, Lefkos, promontori calcarei, 4 July 2000, Brullo & Giusso s.n. (CAT); Karpathos, Ammopi, costa rocciosa calcarenitica, 29 June 2002, Brullo & Giusso s.n. sheet 1 (CAT, Herb. Erben); Karpathos, costa calcarea a Sud di Lefkos, 1 July 2002, Brullo & Giusso s.n. (CAT, Herb. Erben); Karpathos, Kira Panagia, rupi calcaree dolomitiche, 2 July 2002, Brullo & Giusso s.n. (CAT, Herb. Erben); Karpathos, Ahata, 2 July 2002, Brullo & Giusso s.n. (CAT, Herb. Erben); Karpathos, Tal zwischen Aperi und der Achata-Bucht, trockenes Bachbett mit Sandbänken und Felsen, 6 October 2002, Kalheber 02–1690, 02–1693 (Herb. Kalheber); Karpathos, Lefkos, C. Liani Pounda, 22 August 2003, Brullo & Bacchetta s.n,,sheet 1 (CAT, Herb. Erben).

Figures

FIGURE 20. Limonium carpathum.Rechinger fil. A) Habit; B) Spikelet; C) Outer bract; D) Middle bract; E) Inner bract; F) Calyx; G) Calyx lobes; H) Leaves. (Habit and leaves drawn from “Karpathos, Pigadia, Brullo & Guarino s.n., CAT”, spikelet details from “K.H. & F. Rechinger 8241, M”).

FIGURE 92. (Map 7) Geographical distribution of Limonium calliopsium (A); L. carpathum (B); L. chersonesum (C); L. chrisianum (D); L. contractum (E); L. cornarianum (F); L. hierapetrae (G); L. sougiae (H).