Gromphas lemoinei Waterhouse, 1891

Cupello, Mario & Vaz-de-Mello, Fernando Z., 2015, A new species and the phylogeny of the South American genus Gromphas Brullé, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini), Journal of Natural History 50, pp. 943-969 : 954

Gromphas lemoinei Waterhouse, 1891

( Figures 5 – 6 )

Additional material examined

VENEZUELA: TÁCHIRA: Santo Domingo, Sta . Rosa Valz, 13 August 1983, Havranek (?) col. – 1 male ( BMNH) .

Figures

Figure 5. Updated distribution of the six species of Gromphas.

Figure 6. Phylogeny of Gromphas. Above: Cladogram showing the phylogenetic relationship of the species of Gromphas and the evolution of their characters. Black circles indicate uncontroverted synapormorphies; white circles with black margin, controverted synapomorphies; red circles, ambiguous uncontroverted synapomorphies; and white circles with red margin, ambiguous controverted synapomorphies. Below: Support values given for each clade are decay index (above branch) and bootstrap (below branch).