Ariadna maxima ( Nicolet, 1849 )

Giroti, André Marsola & Brescovit, Antonio Domingos, 2018, The taxonomy of the American Ariadna Audouin (Araneae: Synspermiata: Segestriidae), Zootaxa 4400 (1), pp. 1-114: 20-24

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http://doi.org/ 10.11646/zootaxa.4400.1.1

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Ariadna maxima ( Nicolet, 1849 )
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Ariadna maxima ( Nicolet, 1849) 

Figures 9View FIGURE 9, 11View FIGURE 11, 14‒15View FIGURE14View FIGURE 15, 70View FIGURE 70

Dysdera maxima Nicolet, 1849: 341  , pl. 2, figs 6A ‒D (♀ holotype from Santiago [33°26'16"S; 70°39'01"W], Chile, 1845, C. Gay leg., deposited in MNHNP-AR 9068 (4210), examined).— Keyserling (1877: 230).

Dysdera virens Nicolet, 1849: 342  (5♀ syntypes from Santiago, Chile [33°26'16"S; 70°39'01"W], 1845, C. Gay leg., deposited in MNHNP-AR 9071 (4143), examined).— Keyserling (1877: 230).— Simon (1896: 64, Syn.).— Petrunkevitch (1911: 131).

Dysdera incerta Nicolet, 1849: 342  (2♀ syntypes from Santiago, Chile [33°26'16"S; 70°39'01"W], 1845, C. Gay leg., deposited in MNHNP-AR 9072 (4132), examined).— Keyserling (1877: 230).— Simon (1896: 64, Syn.).— Petrunkevitch (1911: 131).

Dysdera coarctata Nicolet, 1849: 344  , pl. 2, figs 7A ‒C (4 immature syntypes from Santiago, Chile [33°26'16"S; 70°39'01"W], 1845, C. Gay leg., deposited in MNHNP-AR 9070 (4145)), examined).— Simon (1864: 106).— Keyserling (1877: 230).— Petrunkevitch (1911: 131).— Bonnet (1956: 1619).— Beatty (1970: 469, Syn.).

Dysdera longipes Nicolet, 1849: 343  , pl. 2, fig. 8 (1♂ 2♀ syntypes from ValdiVia, Chile [39°49'51"S; 73°13'15"W], 1845, C. Gay leg., deposited in MNHNP-AR 9067 (4133), examined). — Simon (1896: 64, Syn.).— Petrunkevitch (1911: 131).

Segestria pusilla Nicolet, 1849: 345  , pl. 2, fig. 9 (Immature holotype from Chile, should be deposited in MNHNP, lost).— Brignoli (1976: 50).— NEW SYNONYMY.

Ariadna  pusilla— Giroti & Brescovit (2011: 63, Transf. from Segestria  , as nomem dubium).

Ariadna  maxima— Simon (1896: 64, Transf. from Dysdera  ; 1902: 11; 1905: 71).—O. P.- Cambridge (1898: 17).— Petrunkevitch (1911: 131).— Bonnet (1955: 734).— Beatty (1970: 469, figs 2, 18‒19, 21).— Grismado (2008: 349, figs 2, 5F ‒H, 8H ‒I, 9A ‒D).— Giroti & Brescovit (2015: 3, figs 9A, 10A).— World Spider Catalog (2018).

Ariadna ashantica Strand, 1916: 54  (1♂ 2♀ syntypes from Ashanti [06°45'N; 01°30'W], Ghana, H. Simon leg, deposited in SMF 2810View Materials, examined.).GoogleMaps  — NEW SYNONYMY.

Ariadna  levii— Grismado (2008: 345, in part, only the ♀ paratype, misidentification).

Synonymy. Giroti & Brescovit (2011) transferred Segestria pusilla  to Ariadna  as nomen dubium, since the unavailability of the holotype and the indefiniton of the type-locality. Considering characters from Nicolet's description, as the carapace reddish brown and light yellow legs, together with the distribution of Ariadna maxima  in Chile, this species is here proposed as a junior synonym of A. maxima  . Grismado (2008) described a female paratype of Ariadna levii  , depicting the female internal genitalia of a non-type specimen. After the examination of those specimens, we detected that the paratype actually is Ariadna maxima  , due to coloration, macrosetae pattern and leg size in relation with body size. The diffuse abdominal pattern presented by the paratype is usual to A. maxima  . On the other hand, the described and depicted genitalia in Grismado (2008: 344, figs 6D ‒E) belongs to an A. levii  specimen.The oddest synonym proposal involves the African species Ariadna ashantica Strand, 1916  . In his work, Strand (1916) described this species based on a male and the cephalothorax and legs of a possible female from Ashanti region, in Ghana. After the examination of the type specimens, we identified them as A. maxima  , based on the morphology of the male palp and metatarsi I. The occurrence of A. maxima  in a locality so far away of the type locality is rather odd, but it could have occurred by anthropic introduction or even human-error handling and labeling the vial in the museum.

Additional material examined. CHILE: III Región de Atacama: Atacama Province : Copiapó, Barranquilla (27°18'41.4"S; 70°30'22.6"W), 2♀ 1imm., 10.XI.2013, A.D. Brescovit et al. leg. ( IBSP 166848View Materials)GoogleMaps  ; Huasco Alto (28°12'34.0"S; 71°05'40.4"W), 2♀ 1imm., 9.XI.2013, same collectors ( IBSP 166847View Materials)GoogleMaps  ; Caleta Chañaral de Aceituno , 18m (29°28'22.1"S; 71°17'33.7"W), 1♂ 6♀ 7imm., 11.XI.2013, same collectors ( IBSP 166850View Materials)GoogleMaps  ; Carrizalillo , 124m (29°37'38.3"S; 71°14'37.5"W), 1♂ 2imm., same date and collectors ( IBSP 166849View Materials)GoogleMaps  ; III Región de Coquimbo: Elqui Province: La Higuera, Los Choros , Llano Choros [29°14'02.4"S; 71°22'02.4"W], 1♀, I.S. E. leg. ( LEULS)GoogleMaps  ; Los Choros [29°15'35.5"S; 71°01'41.4"W], 2♀, 25‒27.VIII.1965, D.N. et al. leg. ( LEULS)GoogleMaps  ; Los Choros, 15km de Punta Choros [29°14'27.5"S; 71°25'44.1"W], 1♀, 3.VI.2005, V. M.R. leg. ( LEULS)GoogleMaps  ; 1♀, 8.X.2005, D.V. leg. (LEULS); 1♀, 8.XI.2005, M.A. leg. (LEULS); Punta Choros [29°14'49''S; 71°27'42''W], 12♀, 25‒27.VIII.2005, D.V. et al. leg. ( LEULS)GoogleMaps  ; 2♀, 26.VIII.2005, J.P.A. leg. (LEULS); 1♀ 3imm., 6‒9.X.2005, D.A.Z. leg. (LEULS); 5♀, 19.XI.2005, D.A.Z. & L.S. leg. (LEULS); La Serena, Punta de Teatinos , 39m (29°29'29.1"S; 71°10'26.5"W), 1♀, 8.XI.2013, A.D. Brescovit et al. leg. ( IBSP 166846View Materials)GoogleMaps  ; Punta de Teatinos [29°55'25.6"S; 71°15'27.3"W], 1♀, 7.XI.2006, L.S. E. leg. ( LEULS)GoogleMaps  ; Las Companias [29°52'37.7"S; 71°14'06.3"W], Llaves , 1♀, 23.VI.2007, L. Cortes leg. ( LEULS)GoogleMaps  ; Compania Baja [29°52'45.7"S; 71°14'55.7"W], 1♂, 26.III.2007, Y. L.P. leg. ( LEULS)GoogleMaps  ; Coquimbo Province: La Pampilla [29°57'32.36"S; 71°21'10.87"W], 1♀, VIII.1977, L. Alvarez leg. ( MHNS 645)GoogleMaps  ; Ovalle, La Alcaparra , 342m (30°17'10.6"S; 71°05'44.7"W), 3imm., 8.XI.2013, A.D. Brescovit et al. leg. ( IBSP 166845View Materials)GoogleMaps  ; Ovalle , 337m (30°20'26.520"S; 71°6'32.580"W), 1♂ 9♀ 8imm., 8.XI.2013, A.D. Brescovit et al. leg. ( IBSP 166844View Materials)GoogleMaps  ; Camino a Fray Jorge , 388m (30°21'27.8"S; 71°23'31.9"W), 3♀ 2imm., 12.XI.2013, A.D.GoogleMaps 

Brescovit & I.L.F. Magalhães leg. (IBSP 166851); Combarbalá, La Ligua Baja (31°09'02.2"S; 71°17'07.8"W), 1♀, 7.XI.2013, A.D. Brescovit et al. leg. (IBSP 166843); Canela Baja, Camino a Canela Alta, 429m (31°13'34.4"S; 71°11'49.7"W), 2♀1imm., same date and collector (IBSP 166842); Canela Baja (31°14'36.8"S; 71°16'33.7"W), 15♀8imm., same date and collector (IBSP 166841); Choapa Province: Illapel, Valle de Pupio, Cuz-Cuz, 204m (31°24'0.07"S; 71°10'33.78"W), 1♂ 3♀ 2imm., 8.IV.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166832); Salamanca, El Tambo, 403m (31°27'46.97"S; 71°02'24.39"W), 2♀ 3imm., same date and collectors (IBSP 166834); Salamanca, 489m (31°28'14.69"S; 70°35'31.24"W), 2♀ 1imm., same date and collectors (IBSP 166833); Los Vilos, near Puerto Punta Chungo, 19m (31°31'33.67"S; 71°17'41.02"W), 1♀ 1imm., same date and collectors (IBSP 166831); 10km S of Los Vilos [31°55'S; 71°31'W], 1♀ 1imm., I.1989, P. Goloboff leg. (MACN-Ar 10099); Los Vilos [31°55'S; 71°31'W], 3imm., 30.IX.1983, E. Maury leg. (MACN-Ar 10098); Ruta 5 km 212, Quebrada 16km N Pichidangui, 10m (32°01'00.5"S; 71°30'29.0"W), 2♂, 20.II.2005, M. Ramírez & F. Labarque leg. (MACN-Ar 10787; 10788); Pichidangui, Valle de Quillimari, 59m (32°03'53.35"S; 71°18'14.07"W), 1♀, 7.IV.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166830); Pichidangui [32°08'S; 71°32'W], 1♂, 12.VIII.1966, E.I. Schlinger leg. (CAS 9046571); Pichidangui (32°08'14.28"S; 71°32'3.33"W), 1♀ 12imm., 20.II.2012, M.J. Ramírez et al. leg. (MACN-Ar 28670); III Región de Valparaíso: Petorca Province: Quebrada de Chivato, 1 km S of Los Molles [32°14'22"S; 71°30'54"W], 1♀, 30.IX.1988, P. Goloboff et al. leg. (MACN-Ar 10106); Papudo, 76m (32°18'11,08"S; 71°16'45,37"W), 5♀ 9imm., 9.IV.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166835); Zapallar, La Laguna, 54m (32°23'22,56"S; 71°15'15,84"W), 15♀4imm. same date and collectors (IBSP 166836; 166837); Pichicuy [32°20'18"S; 71°27'36"W], 20m, 1♀, 9.II.1986, N.I. Platnick leg. (AMNH, paratype of A. levii  ); San Felipe de Aconcagua Province: San Felipe, El Guayacán, 370m (32°17'54,02''S; 70°33'30,02"W), 1♀, 6.IV.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166827); San Felipe, 670m (32°25'24,96"S; 70°26'32,74"W), 2♀, same date and collectors (IBSP 166825); Putaendo, 947m (32°20'24"S; 70°25'33,63"W), 1♂ 3♀, same date and collectors (IBSP 166826); Quillota Province: Quillota, Cuesta El Melón, elev. 520m (32°52'S; 71°15'W), 1♀, 10.I.1985, N.I. Platnick & O.F. Francke leg. (AMNH); Ocoa, Borda do Parque Nacional La Campana, 355m (32°32'59,27"S; 71°03'15,11"W), 1♂ 1♀ 1imm., 23.III.2013, A.D. Brescovit et al. leg. (IBSP 166788); Ocoa, Borda do Parque Nacional La Campana, Palmas de Ocoa, 440m (32°33'36,1"S; 71°03'4,6"W), 2♀ 1imm., same date and collectors (IBSP 166789); Los Andes Province: Los Andes, Juncal, 1950m [33°02'25"S; 70°05'01"W], 1♀, I.1984, P. Goloboff leg. (MACN-Ar 10102); Valparaíso Province: Los Andes, Río Blanco, 1031m (32°30'47.88''S; 70°16'21.756''W), 3♀, 6.IV.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166945); Casablanca [33°19'S; 71°25'W''], 1♀ 2imm. (MNRJ 1094); Juan Fernandez Islands [33°38'42,5"S; 78°49'23,48"W], Masafuera, 2♂ 4♀ 10imm., II.1978 (MHNS 765); 1♀1imm., R. Wachenknechr leg. (MNRJ 588); 1♀, Dr. Plate leg. (NHM 222.); 1♂ (NHM); 3♀2imm., 1905, Porter leg. (MNHNP-AR 7046); RM Región Metropolitana de Santiago: Santiago Province: Las Chilcas, Camino a Llaya Llay, Puente Pedro Bustos M., 616m (32°31'16,32"S; 70°30'31,32"W), 3♀ 1imm., 23.III.2013, A.D. Brescovit et al. leg. (IBSP 166787); Santiago, Caminho para Farillones, 1994m (33°12'41,4"S; 70°11'51,35"W), 2♂ 6♀ 1imm., 21.III.2013, A.D. Brescovit et al. leg. (IBSP 166783); Camino para Farillones, 1262m (33°12'46,43"S; 70°13'18,48"W), 3♂6♀2imm. (IBSP 166782); Valle Lo Campino, 482m (33°13'55,16"S; 70°26'4,84"W), 1♀, 24.III.2013, A.D. Brescovit et al. leg. (IBSP 166790); Cerro San Cristóbal, 650m (33°15'31,60"S; 70°22'45,48"W), 2♂ 15♀ 3imm., same date and collectors (IBSP 166792); Cuesta Lo Prado, 499m (33°16'29,67"S; 70°32'20,97"W), 1♂ 15♀ 5imm., same date and collectors (IBSP 166791); Guayacán, Río Colorado (33°59'S; 70°37'W), 1♀, I.1984, P. Goloboff leg. (MACN-Ar 10100); Cerro Santa Lucía [33°26'16"S; 70°39'01"W], 1♂, II.1961, A.F. Andres leg. (AMNH); Talagante [33°40'S; 70°52'W], 1♀, VII.1961, O. Ramírez leg. (AMNH); Cantillana [33°58'S; 70°58'W], 1♀, 10‒22.XII.1983, L. Irarrazaval leg. (AMNH); Santiago [33°26'16"S; 70°39'01"W], 11♀ 3imm., 20.X.2009, T.H. Kawamoto leg. (IBSP 161292; 166664); 1♀ 1imm., 1899, Silv leg. (MNHNP-AR); 1♂, 25.IV.1985 (MHNS 882); Lo Barnechea [33°22'15"S; 70°26'00"W], Puente El Ñilhue, Camia Tarellones, 1♀, I.1962, Mann leg. (MHNS 602); Cordillera Province: San José de Maipo, Cajon del Maipo, Camiño Central, Hidrelétrica Los Maitenes, 1001m (33°20'32,28"S; 70°12'17,28"W), 3♀ 1imm., 22.III.2013, A.D. Brescovit et al. leg. (IBSP 166784); San José de Maipo, Cajon del Maipo, El Manzano, 880m (33°20'51,72"S; 70°14'48,83"W), 3♂1♀1imm., same date and collectors (IBSP 166786); San José de Maipo, Cajon del Maipo, Pasando San José de Maipo, 1005m (33°24'16,91"S; 70°12'28,79"W), 2♀, same date and collectors, (IBSP 166785); San José de Maipo, Trilha para Refúgio Lagunillas [33°41'S; 70°20'W], 1♀, 3.III.2007, E. Álvares leg. (IBSP 73492); Maipo Province: Buin, Linderos [33°45'46"S; 70°44'07"W], 4♀ 3imm., V.1963, Fritz leg. (MACN- Ar 10119); Huélquen, 434m (33°29'16,79"S; 70°23'17,15"W), 1♂ 2♀ 1imm., 25.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166793); Huélquen, 390m (33°31'29,78"S; 70°24'14,5"W), 1♀ 1imm., same date and collectors (IBSP 166794); VI Región de O'Higgins: Cachapoal Province: Machalí, Coya, 780m (34°07'04"S; 70°22'7,35"W), 4♀ 2imm., 5.IV.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166828); Machalí, Coya, 873m (34°07'17,07"S; 70°18'20,05"W), 4♀, same date and collectors (IBSP 166823; 166824); Machalí, Los Nogales, 653m (34°08'4,95"S; 70°23'44,7"W), 25♀ 5imm., same date and collectors (IBSP 166821; 166822); Doñihue, camino para Doñihue, 449m (34°06'47,44"S; 70°30'25,09"W), 33♀ 4imm., 25.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166795); Cachapoal, R. N. Rio de Los Ciprestes, sector Las Arpas, elev. 1066m (34°16'52,1"S; 70°27'04,2"W), 3♀ 3imm., 20. VII.2 0 10, A. Ojanguren et al. leg. (MACN-Ar 29460; 29461; 29463); Coltauco, Próximo ao Cemitério Municipal, 252m (34°10'7,96"S; 71°03'19,65"W), 5♀ 3imm., 25.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 16 6796); Coltauco, entre Coltauco e Telmo, 220m (34°13'40,36"S; 71°54'0,35"W), 2♂ 9♀ 1imm., 26.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166797); Pichidegua, Pataguas Cerro, 117m (34°11'20,65"S; 71°13'47,31"W), 4♀ 4imm., same date and collectors (IBSP 166798); Pichidegua, 152m (34°12'25,92"S; 71°11'43,04"W), 3♂ 3♀ 4imm., same date and collectors (IBSP 166799); San Vicente [34°26'22"S; 71°04'34"W], 3♀, Silv leg. (MNHNP-AR); Itahue, 241m (35°04'31,79"S; 71°12'59,11"W), 9♀ 7imm., 26.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166 802); Cardenal Caro Province: Litueche, 309m (34°32'56,04"S; 71°24'38,52"W), 14♀ 5imm., 4.IV.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166820); Marchihue, 230m (34°11'54,42"S; 71°21'58,6"W), 6♀ 2imm, same date and collectors (IBSP 166819); Colchagua Province: Palmilla, Isla de Yaquil, 156m (34°20'25,94"S; 71°14'37,75"W), 3♀, 4.IV.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166818); San Fernando, 346m (34°21'53,67"S; 71°00'19,26"W), 32♀8imm., same date and collectors (IBSP 166 816); Puquillay Alto, 338m (34°23'52,62"S; 71°56'51,35"W), 2♂ 12♀ 8imm., same date and collectors (IBSP 166817); Santa Cruz, Quinahue, Pista de Motocross, 185m (34°25'10,84"S; 70°11'49,16"W), 4♀ 2imm., 26.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166800); Chépica, Rincón de los Navarros, 235m (34°29'22,63"S; 71°07'37,09"W), 1♀, same date and collectors, (IBSP 166801); VII Región de Maule: Curicó Province: Curicó [34°59'S; 71°14'W], 10km E Los Queñes, 1000m, 1♀, 23.II ‒9.III.2001, J.E. Barriga leg. (CAS 904 6537); Curicó, Los Niches, Fundo Barriga, elev. 299m (35°03'47,4"S; 71°07'22,3"W), 1♀, 18.VII.2010, A. Ojanguren et al. leg. (MACN-Ar 29458); Talca  Province: Talca  , Cerro La Virgen, 150m (35°15'8,06"S; 71°24'41,61"W), 4♀ 1imm., 27.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166803); Gil de Vilches [35°29'S; 70°58'W], 1♀, 7.I.1989, M. Ramírez leg. (MNHNP-AR 14919); 1♀2imm., 16‒17.I.1984, P. Goloboff leg. (MACN-Ar 10112; 10113); 1♀, 7‒8.I.1984, E. Maury leg. (MACN-Ar 10114); Maule Province: Maule, 178m (35°18'14,97"S; 71°25'18,55"W), 3♀, 27.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166 804); Las Garzas, La Suiza, 653m (35°27'49,28"S; 70°35'50,1"W), 1♂ 10♀ 3imm., 2.II.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166814); Tregualemu [35°57'19"S; 72°44'46"W], 2♀, II.1966, I. Moyano leg. (UCCC); Cauquenes Province: Chanco, Reserva Nacional Federico Albert, 45m (35°26'7,07"S; 72°19'15,96"W), 1♀, 27.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166806); Cauquenes, Reserva Nacional Los Ruiles, 180m (35°30'1,54"S; 72°18'21,38"W), 1♀, same date and collectors (IBSP 166805); Cauquenes, Reserva Nacional Los Ruiles, elev. 174m (35°50'02,6"S; 72°30'35,7"W), 1♀ 1imm., 17.VII.2010, A. Ojanguren et al. leg. (MACN-Ar 29465); Linares Province: Linares, El Canelo [35°51'S; 71°36'W], 1♀ 7imm., X.1963, Fritz leg. (MACN-Ar 10110); VIII Región de Bío-Bío: Ñuble Province: San Fabián de Alico, Fundo El Sauce [36°33'S; 71°33'W], 1♂ 4♀ 7imm., 8‒24.I.1986, L.E. Peña leg. (AMNH); Chillán, Chillán Viejo, 142m (36°22'50,01"S; 72°02'44,91"W), 2♀, 1.IV.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 166811); Pinto, Las Trancas, 1150m (36°32'49,45"S; 71°18'24,8"W), 5♀ 9imm., same date and collectors (IBSP 166812); Chillán, Las Trancas [36°36'21"S; 72°06'13"W], 2♂ 1♀ 3imm., II.1980, L. Peña leg. (AMNH); Concepción Province: Talcahuano [36°43'S; 73°07'W], 2♀, 27.VII.1962, Mendéz leg. (UCCC); Hualpén [36°47'32"S; 73°05'44"W], 1♀, 1975, Tapia leg. (UCCC); Peñuelas [36°58'50''S;72°27'7''W], 1♀, 25.III.1978, D. Gasland leg. (UCCC); Concepción; Río Andalien [36°50'S; 73°03'W], 2♂ 4♀ 13imm., 11.I.1983, G. Muñoz leg. (AMNH); Concepción [36°50'S; 73°03'W], 1♀, 12.VIII.1979, Artigas leg. (UCCC); 1♂ 3♀ 1imm., 8.VIII.1955 (UCCC); 1♂, 14. III.2 0 0 7, L. Garavilla leg. (UCCC); 1♀ (MNRJ 3721); Chiguayante [36°55'S; 73°01'W], 3♀ 2imm., 17.II.2005, C. Solar leg. (UCCC); Coronel, Estación Escuadrón [36°55'59"S; 73°09'00"W], 2♂ 5♀ 3imm., 20.IX.1980, N. Cekalovic leg. (AMNH); Bío-Bío Province: Monte Aguila, 132m (37°02'0,16"S; 72°14'55,39"W), 8♀ 4imm., 29.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 16 6807); Tucapel, Huépil, 234m (37°06'23,86"S; 72°01'57,72"W), 4♀, same date and collectors (IBSP 166808); Tucapel, Polcura, 594m (37°11'6,39"S; 71°22'49,15"W), 3♀ 5imm., same date and collectors (IBSP 166809); IX Región de Araucanía: Cautín Province: Angol [37°48'S; 72°43'W], 1♂2♀1imm., 1950, D.S. Bullock leg. (CAS 9046503); Temuco [38°45'S; 72°40'W], 1♀, 26.VII.1989, C. Salazar leg. (UCCC); Malleco  Province: Collipulli [37°57'S; 72°25'W], 9♀ 14imm., 24.IV.1963, F. Mendéz Z. leg. (UCCC); XIV Región de Los Ríos: Valdivia Province: without specific data, 3♀2imm. (MNHNP-AR); X Región de Los Lagos: Osorno Province: Pucatrihue, 15m (40°19'43,6"S; 73°25'51,31"W), 2♀, 31.III.2013, A.D. Brescovit & I.L.F. Magalhães leg. (IBSP 16 6810); Llanquihue  Province: Puerto Montt [41°28'18"S; 72°56'13"W], 1♂, 12.X.2011, A. Taucare-Rios leg. (IBSP 166780); 1♂, 2012, same collector (IBSP 166781); Without specific locality data: Los Pizios?, 1♀ 1imm., F. Vidala leg. (MNHNP-AR); 1♂ 3♀ 11imm. (MNHNP-AR 7042); 1♂ (MNHNP-AR 7044); 1♂ 51♀ 22imm., X.1894, E. Wangles & Latreile leg. (MNHNP-AR 9073); 2♀ (MNRJ 42166); 7♀ 9imm. (MACN-Ar 10108); 5♀ 5imm. (NHM 90.7.1.483‒490); ARGENTINA: Neuquén Province: Lago Moquehue [38°54'12,5"S; 71°16'42,9"W], 1♀, 10.I.1985, E. Maury & Toth leg. (MACN-Ar 10097).

Diagnosis. Ariadna maxima  is similar to A. araucana  and A. lalen  n. sp. due to the morphology of the male metatarsi I and leg macrosetae pattern of females, but differ by the presence of a male metatarsi I sinuous in lateral view ( Fig. 14C ‒DView FIGURE14), and palps rounded, with the midportion and embolus with similar size ( Fig. 15A ‒BView FIGURE 15); females differ by the anterior receptaculum billobulated ( Figs 9BView FIGURE 9, 15C ‒DView FIGURE 15), with a large median glandular ducts plate and interpulmonary fold with a dorsal median membranous crack ( Fig. 9BView FIGURE 9).

Description. Male ( IBSP 166780). Carapace and chelicerae dark reddish brown ( Fig. 14AView FIGURE14). Endites brownish orange; labium dark brownish orange. Sternum brownish orange with darker margins, and four pair of fainted dark marginal areas ( Fig. 14BView FIGURE14). Palps brownish yellow ( Fig. 15A ‒BView FIGURE 15). Legs brownish orange, with pairs I ‒II darker and with dark pigment forming diffuse rings in the distal region of the metatarsi and tarsi ( Fig. 14C ‒EView FIGURE14). Abdomen grayish, lighter ventrally, and with a diffuse dark dorsal pattern ( Fig. 14A ‒BView FIGURE14). Total length 10.0; carapace 5.2 long, 3.6 wide. Palps: tibia incrassated, with the dorso-ventral diameter equal to the bulb; cymbium with a dorsal chemosensory patch; bulb sub-rectangular, mid-portion slightly shorter than the embolus; embolus thickened, slightly curved, and ending abruptly ( Fig. 15A ‒BView FIGURE 15). Legs: slender and coated with setae ( Fig. 14C ‒EView FIGURE14); tarsal brush of putative chemosensory setae present in legs in pairs I ‒IV, but reduced in legs I, III ‒IV; metatarsus I sinuous, ventrally bent in lateral view ( Fig. 14C ‒DView FIGURE14). Leg formula: II-I-IV-III. Leg measurements: I femur 5.88, patella 2.0, tibia 5.12, metatarsus 5.52, tarsus 1.24, total 19.76; II 5.72, 2.12, 5.4, 5.76, 1.16, 20.16; III 4.28, 1.64, 3.32, 3.6, 0.92, 13.76; IV 4.72, 2.08, 3.96, 3.72, 0.96, 15.44. Macrosetae: I femur D0-0-0-0-0/1-0-2-0, P0-0-0-0-2-0-1-0; tibia P1-0-0-1-0-0-1-1-0, vp1-0-1-0-1/0-0-1-0, vr1/2-0-1-0-1-0-1-0, R1-1-1-1-1-1-1-1-0; metatarsus vp1-1-1-1-0-0-1, vr1-0-0/1-1-0-0-1, r0-1-0-0-0-0-0-0 ( Fig. 14C ‒EView FIGURE14); II femur D0-0-0-0-0/1-1/0-2-0, P0-0-0-0-1-0-1-0; tibia P0-1-0- 1-0-1-0-0, vp1-0-1-0-1-0-1-0, vr2-0-1-0-1-0-1-0, R1-1-1/0-1-1-1-1-1-1-0; metatarsus vp1-1-1-1-0-0-1, vr1-0-0/1- 1-0-0-1, r0-1-0-0-0-0-0-0; IV femur D2/1-1-1/2-1/2-0/1-1/0-1/2-1/2-1/2-1-0-0, R0-0-0-0-0-0-0-0-0-1-0-0; tibia vp0-0-0-0-0-0-1, vr0-0-0-0-0-0-1; metatarsus vp0-0-0-0-0-0-2, vr metatarsal comb with 4 macrosetae, r0-0/1-0-0- 0-0-0.

Female ( IBSP 161292). Carapace and chelicerae dark reddish brown, almost black ( Fig. 14FView FIGURE14). Endites brownish orange; labium dark brownish orange. Sternum brownish orange with darker margins, and four pairs dark marginal areas ( Fig. 14GView FIGURE14). Palps brownish orange, darkening gradually in distal direction. Legs brownish orange, with pairs I ‒II darker and with dark pigment forming diffuse rings in the distal region of metatarsi and tarsi ( Fig. 14H ‒IView FIGURE14). Abdomen as in male ( Fig. 14F ‒GView FIGURE14). Total length 14.24; carapace 5.96 long, 3.72 wide. Legs: robust and densely coated with setae; legs I ‒II with ventro-prolateral and ventro-retrolateral tibial macrosetae of similar size, and metatarsal macrosetae of different size ( Fig. 14H ‒IView FIGURE14). Leg formula: I-II-IV-III. Leg measurements: I femur 4.4, patella 1.92, tibia 3.6, metatarsus 3.32, tarsus 0.88, total 14.12; II 4.08, 2.0, 3.4, 3.16, 0.76, 13.4; III 3.24, 1.68, 2.32, 2.32, 0.76, 10.32; IV 4.12, 2.04, 3.36, 2.72, 0.8, 13.04. Macrosetae: I, femur D0-0-0-0-0-2-0, P0-0-0-0-2-1-0; tibia P0-1-0-1-0-1-0, vp1-0-1-0-1-0-1, vr1-0-1-0-1-0-1, R0-1-0-1-0-0/1-0; metatarsus vp1-1-1-1-1-0/1-1-1-1-1-1, vr1-1-1-1-1-1-1-1-1-1-1 ( Fig. 14H ‒IView FIGURE14); II, femur D0-0-0-0-0-1/2-0, P0-0-0-0-1-1-0; tibia P0-1-0-1-0-1-0, vp1-0-1- 0-1-0-1, vr1-0-1-0-1-0-1, R0-1-0-1-0-1/0-0; metatarsus vp1-1-1-1-1/0-1-1-1-1-1-1, vr1-1-1-1-1/0-1-1/0-1-1-1-1; IV, metatarsus vp0-0-0-0-0-0-1, metatarsal comb with 4 macrosetae. Internal genitalia: anterior receptaculum with dorsal and ventral lobe with similar length, and glandular plate located in the median region of the receptaculum and proximal regions of the lobes ( Figs 9BView FIGURE 9, 15C ‒DView FIGURE 15).

Variation. Male (n=10): Total length 7.7‒11.15; carapace 3.9‒5.6 long, 2.7‒3.8 wide; femur I 3.7‒5.88. Female (n=10): Total length 10.43‒20.25; carapace 5.25‒7.75 long, 3.31‒5.0 wide; femur I 4.18‒5.56.

Distribution. Central and south of Chile, Argentina (Neuquén Province), and Ghana (Ashanti region) ( Fig. 70View FIGURE 70).

IBSP

Instituto Biologico de Sao Paulo

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Segestriidae

Genus

Ariadna

Loc

Ariadna maxima ( Nicolet, 1849 )

Giroti, André Marsola & Brescovit, Antonio Domingos 2018
2018
Loc

Ariadna ashantica

Strand 1916: 54
1916
Loc

Dysdera maxima Nicolet, 1849 : 341

NiColet 1849: 341
1849
Loc

Dysdera virens Nicolet, 1849 : 342

NiColet 1849: 342
1849
Loc

Dysdera incerta Nicolet, 1849 : 342

NiColet 1849: 342
1849
Loc

Dysdera coarctata

Nicolet 1849: 344
1849
Loc

Dysdera longipes Nicolet, 1849 : 343

NiColet 1849: 343
1849
Loc

Segestria pusilla

Nicolet 1849: 345
1849
Loc

Ariadna

Audouin 1826
1826
Loc

Ariadna

Audouin 1826
1826
Loc

Ariadna

Audouin 1826
1826
Loc

Segestria

Latreille 1804
1804
Loc

Dysdera

Latreille 1804
1804