Bettotania C. Willemse, 1933

Storozhenko, Sergey Yu., 2021, On the exact position of the tribe Incolacridini in the modern classification of grasshoppers (Orthoptera: Acrididae), Zootaxa 4970 (1), pp. 106-118 : 111

Genus Bettotania C. Willemse, 1933

Bettotania C. Willemse, 1933: 75 (type species: Bettotania maculata C. Willemse, 1933 , by original designation); C. Willemse, 1956: 112; Ingrisch, 1989: 227; Otte, 1995: 277; Yin et al., 1996: 96.

Paracelebesia Miller, 1935a: 697 (type species: Paracelebesia festiva Miller, 1935 , by original designation); synonymized with Bettotania by C. Willemse (1956).

Description. Head in frontal view oval; frontal ridge narrow, almost reaching the clypeus. Face in lateral view reclinated; frontal ridge considerably projecting between the antennae and forming a widened rostrum ( Fig. 1 ). Ventral genicular lobes of the hind femora narrowly triangular with apex obtuse, without spine. Male 10th abdominal tergite with furculae. Male supra-anal plate long-oval, with a median basal impression, lateral margins with a pointed tooth somewhat behind the middle, apex rounded. Male cerci compressed, conical, apex pointed, inner side without tooth. Valves of ovipositor relatively long, with apex rounded and crenulate ( Fig. 1 ). Male genitalia: epiphallus divided and asymmetrical; bridge relatively broad; left lophus long, finger-shaped; right lophus small, rounded; ancorae absent; oval sclerites attached to epiphallus ( Fig. 19 ); apodemes broad; valves of cingulum short; basal valves of penis connected with apical ones by a sinuate, long, unbroken flexure ( Fig. 21 ).

Species included. The genus consists of four species distributed in Malaysia (Peninsular Malaysia and Borneo Island).

Figures

FIGURES 1–14. Subfamily Incolacridinae. 1. Bettotania cincifemur, female body, lateral view (schematic after Miller, 1935); 2. Stolzia rubromaculata, male head and pronotum, lateral view (schematic after C. Willemse, 1930); 3. Stolzia javana, female head, frontal view; 4–7. Incolacris rubritarsi, female: 4. Head and pronotum, lateral view; 5. Head, frontal view; 6. Apex of abdomen, lateral view (after Hollis, 1975); 7. The same, ventral view (after Hollis, 1975); 8–12. Incolacris jianfengensis comb. nov., male (after Zheng & Ma, 1989): 8. Head and pronotum, lateral view; 9. The same, dorsal view; 10. Apex of abdomen, dorsal view; 11. The same, lateral view; 12. Hind knee, lateral view; 13. Asymmetritania asymmetrica comb. nov., male apex of abdomen, dorsal view (after Ingrisch, 1989); 14. Asymmetritania vietnamensis comb. nov., male hind knee, lateral view.

FIGURES 15–24. Male genitalia of the subfamily Incolacridinae (15–18 after Hollis, 1975 and 19–24 after Ingrisch, 1989). 15. Incolacris trifasciata, epiphallus, dorsal view; 16. The same, Stolzia javana; 17. The same, Stolzia aberrans; 18. The same, Stolzia fasciata; 19–21. Bettotania maculata: 19. Epiphallus, dorsal view; 20. Phallic complex (epiphallus removed), dorsal view; 21. The same, lateral view; 22–24. Asymmetritania asymmetrica comb. nov.: 22. Epiphallus, dorsal view; 23. Phallic complex (epiphallus removed), lateral view; 24. The same, dorsal view. (Ae—ancorae; An—anterior projections of epiphallus; Ap—apical valves of penis; Apd—apodemes; Bp—basal valves of penis; Br—bridge; Cv—valves of cingulum; Fx—flexure; Lel—left lophi of epiphallus; Ler—right lophi of epiphallus; Os—oval sclerites; Rm—rami of cingulum; Sh—ectophallic sheath of penis; Zyg—zygoma).