Creophilus lanio, (ERICHSON)
publication ID |
https://doi.org/ 10.1111/j.1096-3642.2011.00725.x |
publication LSID |
lsid:zoobank.org:pub:FBFE9195-BE04-4AFE-9417-6E38BCE6AB84 |
persistent identifier |
https://treatment.plazi.org/id/039B414F-1941-FFBB-FC41-F9894B5BFD87 |
treatment provided by |
Valdenar |
scientific name |
Creophilus lanio |
status |
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8. CREOPHILUS LANIO (ERICHSON) View in CoL
( FIGS 2H–J, 3Q, 4J View Figure 4 , 7B View Figure 7 , 29 View Figure 29 , 30 View Figure 30 )
Staphylinus lanio Erichson, 1839: 352 View in CoL . Type locality: ‘Terra Van-Diemenii’; Gravenhorst, 1806: 126.
Staphylinus oculatus View in CoL var. Gravenhorst; Dejean, 1821: 21; Erichson, 1839: 352 (synonym of C. lanio View in CoL ).
Creophilus lanio View in CoL ; Fauvel, 1875: 56; Bernhauer & Schubert, 1914: 398; Lea, 1925: 229 (synonym of erythrocephalus View in CoL ); Steel, 1949: 58, figs 2, 5, 7 and 8; Radford, 1981: 174 (as C. lania , error for lanio View in CoL ); Herman, 2001b: 3317.
Emus lanio ; Fauvel, 1877: 250; Fauvel, 1878a: 248; Fauvel, 1878b: 541.
Creophilus erythrocephalus var. lanio View in CoL ; Olliff, 1887: 492–493.
Type material: Staphylinus lanio Erichson. Lectotype (here designated). ♀, ‘[orange] Type/ 5886/ [grey] lanio| Er.| terra van Diem [Tasmania], Sch/ [lilac] Hist.-Coll. ( Coleoptera )| Nr. 5886| Staphylinus lanio Erichs. | Terra v. Diem., Schayer| Zool. Mus. Berlin / [red] SYNTYPUS | Staphylinus | lanio Erichson, 1839 | labeled by MNHUB 2004/ FMNH-INS 0000 016 774/ [red] LECTOTYPE | Staphylinus | lanio Erichson, 1839 | designated by| D. J. Clarke 2008’ (in ZMHB). Specimen missing left mesotarsus. Paralectotypes (3). All with labels ‘[orange] Type/ [lilac] Hist.-Coll. ( Coleoptera )| Nr. 5886| Staphylinus lanio Erichs. | Terra v. Diem., Schayer| Zool. Mus. Berlin / [red] SYNTYPUS | Staphylinus | lanio Erichson, 1839 | labeled by MNHUB 2004/ [yellow] PARA- LECTOTYPE | Staphylinus | lanio Erichson, 1839 | designated by| D. J. Clarke 2008’: 2♂, ‘ FMNH-INS 0000 016 775’; ‘ FMNH-INS 0000 016 777’; 1♀, ‘ FMNH-INS 0000 016 776’ (in ZMHB).
Other material examined: 2743 specimens. See supporting information, Appendix S 1.
Diagnosis: With characters of the erythrocephalus - group; head orange-red, with large diffuse subquadrate or cordiform black spot ( Fig. 7B View Figure 7 ); right mandible with three teeth ( Fig. 29A View Figure 29 ); elytra black, humeri glabrous, conspicuously sculptured; abdominal segment IX orangeish-brown; parasutural 1 and humeral macrosetae absent, tergal chaetotaxic formula = 4-6-6–6-4-6.
Description: Measurements ( N = 10♂, 10♀). Forebody length: ♂ 5.5–9.8 mm, ♀ 5.3–7.8 mm. See supporting Table S 5 for comparison of ranges of male and female ratios. Head. Head orange-red, narrowly black around mouthparts and antennal fossae, with large diffuse irregularly subquadrate or cordiform black spot ( Fig. 7B View Figure 7 ), usually concealing dorsal tentorial pits, frequently with midlongitudinal line of black pigment continuing to frontoclypeal margin and to neck; strongly trapezoidal, much wider posteriorly; HW/ HL = 1.35–1.59; shining, without distinct microsculpture; eyes small to moderately large ( EYL / HL = 0.38–0.56), dorsolateral, lateral margins of head visible in dorsal view (not obscured by eye); HL 1/ HL 2 greater in females than males (♂ = 1.21–1.83, ♀ = 1.63–2.00); antennae as in Figure 29B View Figure 29 , antennomeres 1–6 black, 7–11 greyish-black, 11 as long or very slightly longer than 9–10 together; mandibles as in Figure 29A View Figure 29 , moderately longer than head in large males, subequal to head in females ( ML / HL ♂ = 0.81– 1.32, ♀ = 0.82–1.05), right mandible with three teeth, T 3 largest. Thorax and abdomen. Pronotum ( Fig. 29F View Figure 29 ) slightly transverse ( PW / PL = 1.09–1.25); PL 1.22–1.45 ¥ ESL; with basolateral margins distinctly emarginate, hind angles distinct; with sparse peripheral setae and short, sparse vestiture on lower anterolateral declivities; elytra uniformly black, humeral regions shining, distinctly callused, disc densely setose, sparsely rugosely sculptured, especially on humeri; wings fully developed, black with distinct black spot in medial field between MP 3 and MP 4 veins; abdomen shining black, except for lightly pigmented orangeish-brown ninth segment; tergite VII with well-developed palisade fringe; styli bicoloured, orangeish-brown basally and black distally. Male genitalia and secondary sexual characters. Ventrolateral carina of large males indistinct, partially obliterated by punctures and secondary anastomosing ridges. Aedeagus as in Figure 29E View Figure 29 ; median lobe apex extended into long blunt point ( Fig. 29I View Figure 29 ), produced slightly dorsally at tip, with paired apicolateral sclerites (as) separated from sclerotized median lobe by distinct membranous strip. Paramere as in Figure 29D View Figure 29 . Internal sac inverted as in Figure 29E View Figure 29 ; ventral sclerite (vs) small, wider than long, distinctly notched apically ( Fig. 25G View Figure 25 ). Female internal genitalia. Internal female genitalia as in Figure 29C View Figure 29 ; vaginal plate with paired lateral sclerites (pls), posterolateral areas membranous; vaginal fold forming large finely rugose sclerite ( Fig. 29C View Figure 29 , vf). Chaetotaxy. Elytra without parasutural 1 and humeral macrosetae; elytral discal series with 3–4 macrosetae; metaventrital macroseta absent or undetected; tergal chaetotaxic formula = 4-6-6-6-4-6, medial macrosetae absent on tergite III, inner lateral macrosetae absent on tergite VII; second gonocoxal macroseta absent.
Variation: The dorsomedial spot in C. lanio is very variable. Occasionally, usually in smaller specimens, it is expanded to cover most of the area between the eyes, although always surrounded by the usual orange coloration of the head.
Comparison: Creophilus lanio may be immediately distinguished from C. erythrocephalus and C. imitator by the diffuse, never circular cranial spot and by the lightly pigmented ninth abdominal segment. Contrary to Steel (1949: 58), all specimens I examined had uniformly shining black elytra, none with a ‘purplish reflection’. Although Cameron (1952: 255) described the elytral sculpturing of C. imitator as ‘less rugose than in lanio ’ it is more distinctive in C. imitator .
Distribution ( Fig. 30 View Figure 30 ): Australia: New South Wales, Tasmania, Victoria, southern Queensland. One record from Western Australia (no locality given, possibly mislabelled).
Biology and ecology: Creophilus lanio is known mostly from forested areas, with few records from pasture and coastal habitats. Most collections are from carrion-baited pitfall traps. Specimens have also been taken at light, in yellow pan traps, in Malaise and window traps, and in dung of various sorts. Habitat: drier woodlands to temperate rainforests of many kinds (e.g. Acacia , Banksia , Eucalyptus , Nothofagus , and Pomaderris ). Altitude: sea level to 1650 m. Phenology: throughout the year. Other biology and lifehistory characteristics are unknown. Larvae and pupae are unknown.
J |
University of the Witwatersrand |
S |
Department of Botany, Swedish Museum of Natural History |
N |
Nanjing University |
HL |
Houghton Lake Wildlife Research Station |
ML |
Musee de Lectoure |
T |
Tavera, Department of Geology and Geophysics |
PW |
Paleontological Collections |
PL |
Západoceské muzeum v Plzni |
MP |
Mohonk Preserve, Inc. |
I |
"Alexandru Ioan Cuza" University |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Genus |
Creophilus lanio
Clarke, Dave J. 2011 |
Creophilus erythrocephalus var. lanio
Olliff AS 1887: 492 |
Emus lanio
Fauvel A 1878: 248 |
Fauvel A 1878: 541 |
Fauvel A 1877: 250 |
Creophilus lanio
Herman LH 2001: 3317 |
Radford WPK 1981: 174 |
Steel WO 1949: 58 |
Lea AM 1925: 229 |
Bernhauer M & Schubert K 1914: 398 |
Fauvel A 1875: 56 |
Staphylinus lanio
Erichson WF 1839: 352 |
Gravenhorst JLC 1806: 126 |
Staphylinus oculatus
Erichson WF 1839: 352 |
Dejean PFMA 1821: 21 |