Teloganodes tristis ( Hagen, 1858 )

Sartori, Michel, Peters, Janice G. & Hubbard, Michael D., 2008, A revision of Oriental Teloganodidae (Insecta, Ephemeroptera, Ephemerelloidea), Zootaxa 1957 (1), pp. 1-51 : 9-11

Teloganodes tristis ( Hagen, 1858)

( Figs. 1–3 , 102 )

Cloë tristis Hagen, 1858:476 .

Leptophlebia tristis ; Eaton, 1873:394.

Teloganodes tristis ; Eaton, 1882:208; 1884:135.

Teloganodes major Eaton, 1884:136 syn. nov.

nec Teloganodes tristis sensu Ulmer 1924:44 ; 1939:512; 627 and subsequent authors Material examined. Besides the type specimens mentioned above (MCZ), very few specimens identified as T. tristis are known. In the collections of BMNH are housed 3 specimens identified as T. tristis , presumably belonging to R. McLachlan’s collection. We have no evidence that these specimens have been seen or studied by Eaton, and none of them (1 male imago, 1 female imago and 1 female subimago) fit Hagen’s specimens diagnosis. In the BMNH collection is housed the lectotypus of Teloganodes major Eaton , a female subimago ( Kimmins 1960). Two syntypes are also held in the collection of the Museum of Comparative Zoology in Cambridge and these specimens were also available for study. One of them is completely embedded in fungi. The other one is a male subimago in good condition. This specimen has also been relaxed, removed from the pin, and stored in ethanol.

Male subimago. Body length: 7.5 mm; forewing length: 11.0 mm; hindwing length: 1.5 mm.

The upper portion of the eye is reddish brown. All characters of wings ( Fig. 2 ) and abdomen are similar to those of the T. tristis lectotypus, except that vein MP2 is attached, not free and crossveins are more numerous (see also Figs 9 and 11 ). The gonopods are three-segmented, the third segment is much shorter than the preceding segment. The styliger plate is definitively convex. Penis lobes are fused for their entire length, except at the apex where a ventral ridge is visible ( Fig. 5 ).

This species has been described from material from the same locality as T. tristis . Consequently, we establish that T. major Eaton, 1884 is a junior subjective synonym of T. tristis ( Hagen, 1858) .

Note on the type locality. In the original description, Hagen (1858) mentioned “Rainbodde” as the type locality. In a subsequent publication ( Hagen, 1859), this name was corrected to Rambodde, due probably to confusion resulting from interpretation of the handwritten label of the collector, Mr. Nietner, a German entomologist who spent some years in Ceylon ( Tennent 1860, p. 247). Rambodde is presently called Ramboda and is a village situated on the road from Kandy to Nuwara Eliya. The exact location of Hagen's “Rambodde” is not known with certainty, but in the 19 th century, there was no village, only tea plantations. The altitude ranges between 1200 m (village Ramboda) and 2000 m (Ramboda Pass)

Known stages. Teloganodes tristis is known only from females and male subimago. The male imago and the nymphal stage remain to be described (but see below).

Figures

FIGURES 1–6. Teloganodes spp. Fig. 1: Forewing of the female subimago lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 2: Forewing of a male subimago syntype of Teloganodes major Eaton [MCZ]. Fig. 3: Hindwing of the female lectotype of Teloganodes tristis (Hagen) [MCZ]; Fig. 4: Hindwing of the male holotype of Teloganodes dentatus Navás [MNHN]; Fig. 5: Genitalia of a male subimago syntype of Teloganodes major Eaton [MCZ]; Fig. 6: Genitalia in ventral view of the male holotype of T. dentatus Navás [MNHN].

FIGURES 98–102. Egg structure of Teloganodes hubbardi (98–101, SEM pictures) and T. tristis (102, light microscope). Fig. 98: Shape of the egg; Fig. 99: Detail of the polar cap; Fig. 100: Micropyle area and crown of sets; Fig. 101: Detail of the uncoiled tread of the epithema; Fig. 102: Partial view of polar caps where crown and uncoiled threads are visible.

FIGURE 9–13. Dudgeodes spp. Fig. 9: Female subimago forewing of Dudgeodes hutanis; Fig. 10: Hindwing of D. hutanis; Fig. 11: Male imago forewing of Dudgeodes pescadori; Fig. 12: Hindwing of D. pescadori; Fig. 13: Genitalia in ventral view of D. pescadori.