Andes Stål, 1866

Wang, Xiao-Ya, Zhi, Yan & Chen, Xiang-Sheng, 2022, Five new species of the genus Andes Stål, 1866 from China (Hemiptera, Fulgoromorpha, Cixiidae), European Journal of Taxonomy 831 (1), pp. 45-66 : 46-48

Genus Andes Stål, 1866

Andes Stål, 1866: 166 .

Leirioessa Kirkaldy, 1907: 112 (synoymized in Muir 1925: 201).

Andes – Muir 1925: 201. — Zhou et al. 1985: 23. — Tsaur et al. 1991: 67.

Type species

Andes undulatus Stål, 1870 (subsequent designation in Muir 1925).

Diagnosis (modified from Tsaur et al. 1991)

Vertex narrowest and transverse at apex, widening to base, with U- or V-shaped basal emargination; lateral carinae strongly elevated; median carina absent. Forewing in resting position steeply tectiform; ScP +RA, RP and MP arising separately from a common point on basal cell or forming a minute common stem. Fore coxa with outer margin straight, subparallel with inner margin, not produced. Hind tibiae without or with several very small lateral spines. Chaetotaxy of hindtarsi 7–9/7–8.

Distribution

Palaearctic Region, Oriental Region, Australian Region and Afrotropical Region.

Checklist and distribution of Chinese species of Andes Stål, 1866

A. balteiformis Wang, Zhi & Chen sp. nov.; China (Guizhou, Guangxi, Fujian).

A. bifidus Wang, Zhi & Chen sp. nov.; China (Yunnan).

A. formosanus ( Matsumura, 1914) ; China (Taiwan).

A. furcutus Wang, Zhang & Chen sp. nov.; China (Guizhou).

A. lachesis Fennah, 1956 ; China (Zhejiang).

A. latanalus Wang & Chen sp. nov.; China (Guangxi).

A. luzonensis Tsaur & Hsu, 1991 ; China (Taiwan).

A. marmoratus ( Uhler, 1896) ; China (Beijing), Japan (Hokkaido, Honshu, Kyushu and Shikoku). A. noctua Fennah, 1956 ; China (Hubei).

A. notatus Tsaur & Hsu, 1991 ; China (Taiwan).

A. othrepte Fennah, 1956 ; China ( Hongkong).

A. pallidus Wang & Chen sp. nov.; China (Guizhou).

A. uncinatus Fennah, 1956 ; China (Guangdong).

Key to males of Chinese species of Andes Stål, 1866

1. Forewing with two eye-like spots of different sizes ( Tsaur et al. 1991: fig. 36a) ............................... ........................................................................................................ A. luzonensis Tsaur & Hsu, 1991

– Forewing without such markings......................................................................................................2

2. Frons and lateral margin of vertex with alternating transverse stripes.............................................3

– Frons and lateral margin of vertex without alternating transverse stripes.......................................5

3. Tegmen with clavus smoky brown at base; tegminal apex near apical angle distinctly dark brown ( Ishihara 1957: fig. 1t) ........................................................................... A. marmoratus ( Uhler, 1896)

– Tegmen without above spots.............................................................................................................4

4. Medioventral process of pygofer triangular; periandrium with a long, strongly sinuate spine, directed cephalad ( Fennah 1956: fig. 1h) ...................................................... A. noctua Fennah, 1956

– Medioventral process of pygofer semicircular; periandrium with a long, slender, C-shaped spine, directed ventrad; female with reduced wax secreting plates ( Tsaur et al. 1991: fig. 34g –i, l) ....... ..................................................................................................... A. formosanus ( Matsumura, 1914)

5. Periandrium with a long, slender, S-shaped spine with its tip directed dorsad ( Tsaur et al. 1991: fig. 35h–i) ............................................................................................. A. notatus Tsaur & Hsu, 1991

– Periandrium without such a spine....................................................................................................6

6. Medioventral process of pygofer at base with a short spine on each side .................................... ........................................................................................................ A. unicinatus Fennah, 1956

– Medioventral process of pygofer without spine..............................................................................7

7. Apical margin of gonostyli deeply emarginate, forming two processes ( Fig. 3G, I ) .................... ........................................................................................... A. bifidus Wang, Zhi & Chen sp. nov.

– Gonostyli not forming two processes..............................................................................................8

8. Periandrium on left laterally with a sclerotized, belt-shaped process arising ventrally near apex ( Fig. 1J–M ) ..................................................... A. balteiformis Wang, Zhi & Chen sp. nov.

– Periandrium without a belt-shaped process....................................................................................9

9. Anal segment wider than long in dorsal view ( Fig. 7H ) .... A. latanalus Wang & Chen sp. nov.

– Anal segment longer than wide in dorsal view...........................................................................10

10. Ventral margin of periandrium apically with a bifurcated spinose process ( Fig. 9J–M ) .............. ..................................................................................................... A. pallidus Wang & Chen sp. nov.

– Periandrium without a bifurcate process.......................................................................................11

11. Dorsal margin of periandrium with a short laminal process at about midlength, basal part wide and apical margin bifurcated ( Fig. 5J–M ) ......... A. furcutus Wang, Zhang & Chen sp. nov.

– Periandrium without such a process..............................................................................................12

12. Endosoma of aedeagus basally with two spinose processes ( Fennah 1956: fig. 1e) ...................... .................................................................................................................... A. othrepte Fennah, 1956

– Endosoma of aedeagus basally without spinose processes; endosoma coarsely shagreen or subfimbriate ( Fennah 1956: fig. 2m) ............................................................. A. lachesis Fennah, 1956

Figures

Fig. 3. Andes bifidus Wang, Zhi & Chen sp. nOv., ♂. A. Dorsal view. B. Lateral view. C. Head and thorax, dorsal view. D. Face, ventral view. E. Forewing. F. Genitalia, lateral view. G. Pygofer and gonostyli, ventral view. H. Anal segment, dorsal view. I. Gonostyli, lateral view. J. Aedeagus, right side. K. Aedeagus, left side. L. Aedeagus, dorsal view. M. Aedeagus, ventral view. Scale bars: A–D, F–M = 0.5 mm; E = 1.0 mm.

Fig. 1. Andes balteiformis Wang, Zhi & Chen sp. nOv., ♂. A. Dorsal view. B. Lateral view. C. Head and thorax, dorsal view. D. Face, ventral view. E. Forewing. F. Genitalia, lateral view. G. Pygofer and gonostyli, ventral view. H. Anal segment, dorsal view. I. Gonostyli, lateral view. J. Aedeagus, right side. K. Aedeagus, left side. L. Aedeagus, dorsal view. M. Aedeagus, ventral view. Scale bars: A–D, F–M = 0.5 mm; E = 1.0 mm.

Fig. 7. Andes latanalus Wang & Chen sp. nOv., ♂. A. Dorsal view. B. Lateral view. C. Head and thorax, dorsal view. D. Face, ventral view. E. Forewing. F. Genitalia, lateral view. G. Pygofer and gonostyli, ventral view. H. Anal segment, dorsal view. I. Gonostyli, lateral view. J. Aedeagus, right side. K. Aedeagus, left side. L. Aedeagus, dorsal view. M. Aedeagus, ventral view. Scale bars: A–D, F–M = 0.5 mm; E = 1.0 mm.

Fig. 9. Andes pallidus Wang & Chen sp. nOv., ♂. A. Dorsal view. B. Lateral view. C. Head and thorax, dorsal view. D. Face, ventral view. E. Forewing. F. Genitalia, lateral view. G. Pygofer and gonostyli, ventral view. H. Anal segment, dorsal view. I. Gonostyli, lateral view. J. Aedeagus, right side. K. Aedeagus, left side. L. Aedeagus, dorsal view. M. Aedeagus, ventral view. Scale bars: A–D, F–M = 0.5 mm; E = 1.0 mm.

Fig. 5. Andes furcutus Wang, Zhang & Chen sp. nOv., ♂. A. Dorsal view. B. Lateral view. C. Head and thorax, dorsal view. D. Face, ventral view. E. Forewing. F. Genitalia, lateral view. G. Pygofer and gonostyli, ventral view. H. Anal segment, dorsal view. I. Gonostyli, lateral view. J. Aedeagus, right side. K. Aedeagus, left side. L. Aedeagus, dorsal view. M. Aedeagus, ventral view. Scale bars: A–D, F–M = 0.5 mm; E = 1.0 mm.