Teruliini, Nielson, 1979

Nielson, M. W., 2011, Tribe Teruliini: new genera and new species of Neotropical leafhoppers with a revised key to genera and species, new records and notes on distribution and taxonomy (Hemiptera: Cicadellidae: Coelidiinae), Zootaxa 2795 (1), pp. 1-45 : 4-6

Key to genera of Teruliini (males)

1. Crown with lateral margins distinctly carinate, anterior margin angulate, distinctly produced beyond anterior margin of eyes. ................................................................................................... 2

- Crown with lateral margins not carinate, anterior margin usually rounded, rarely angulate, usually produced slightly beyond or even with anterior margin of eyes........................................................................ 8

2. Styles symmetrical.................................................................................... 3

- Styles asymmetrical..................................................................... Kravilidius Nielson

3. Style with lateral teeth or setae.......................................................................... 4

- Style glabrous......................................................................................... 5

4. Style with numerous setae at apical ¼; aedeagus long, very slender, with a few long to numerous short spines at apical ½......................................................................................... Carinolidia Nielson

- Style with numerous lateral teeth at apical ½; aedeagus long, robust with numerous long spines throughout length of shaft.......................................................................................... Vernobia Nielson

5. Medium size, slender species, length less than 8 mm; aedeagus tubular or compressed laterally....................... 6

- Large, robust species, length over 9 mm; aedeagus tubular............................................. Terulia Stål

6. Aedeagus tubular with several medial spines along shaft and or subapically....................................... 7

- Aedeagus compressed laterally with single subapical spine...................................... Sprundigia Nielson

7. Pygofer scleriorate on middle of caudal margin; aedeagus nearly straight in lateral view.............. Racinolidia Nielson

- Pygofer not scleriorate; aedeagus with base and apex distinctly curved in lateral view............. Paracarinolidia Nielson

8. Aedeagus symmetrical or nearly so....................................................................... 9

- Aedeagus distinctly asymmetrical........................................................................ 10

9. Mesonotum small, median length equal to median length of pronotum; aedeagus with dorsobasal flange and paired subapical ventral spines.......................................................................... Laevilidia Nielson

- Mesonotum large, median length greater than median length of pronotum; aedeagus without such processes.................................................................................................... Panolidia Nielson

10. Styles symmetrical; aedeagus single..................................................................... 11

- Styles symmetrical; aedeagus bipendulate.................................................... Harasupia Nielson

11. Subgenital plate with few to numerous fine hairlike setae on surface or along margin, apex usually rounded............. 13

- Subgenital plate with numerous, very long, coarse setae on margin a apical ½, apex usually pointed with distinct spine apically Labocurtidia Nielson

12. Subgential plate with apical half elliptical................................................................. 14

- Subgenital plate with apical half ligulate ( Fig. 2 )........................................... Freytagolidia gen. nov.

13. Style very short, extending to or slightly beyond base of aedeagus in dorsal view................................. 15

- Style long, extending to or beyond midlength of aedeagus in dorsal view......................................... 16

14. Aedeagus with subapical process directed distally............................................... Conbalia Nielson

- Aedeagus with subapical process directed basally................................................ Yochlia Nielson

15. Aedeagus long, nearly as long as or longer than style........................................................ 17

- Aedeagus very short, about half as long as style.............................................. Derriblocera Nielson

16. Pygofer in lateral view with 1–2 caudal processes, processes small, lobe-like, short or long and narrow or broadly digitate, with or without tuft of setae on caudal margin.................................................................. 24

- Pygofer in lateral view usually without such processes, usually with tuft of setae on caudal margin.................... 18

17. Style glabrous....................................................................................... 19

- Style usually densely setaceous distally....................................................... Perulidia Nielson

18. Aedeagus with gonopore subapical, setaceous subapically, apically or glabrous; distribution Neogeic or South America.. 20

- Aedeagus with gonopore apical, setaceous medially; distribution Mexico........................... Mexolidia Nielson

19. Style as long as or much longer than aedeagus, apophysis robust.............................................. 21

- Style shorter than aedeagus, apophysis slender............................................................. 22

20. Aedeagus with numerous fine, short setae subapically; style much longer than aedeagus.............. Cochanga Nielson

- Aedeagus with 2–3 very long, robust setae, sometimes with short, stout spines; style about as long as aedeagus.................................................................................................. Bolidiana Nielson

21. Head narrower than pronotum; subgenital plate with numerous stout spines along outer lateral margin; aedeagus broadly tubular, nearly always with several subapical, short to long setae.................................................. 23

- Head nearly as wide as pronotum; subgenital plate with numerous, very fine setae in distal 1/3; aedeagus narrowly tubular, with single spine apically.................................................................. Licolidia Nielson

22. Aedeagus with shaft somewhat compressed laterally, about twice as broad in lateral view than in dorsal view, with a tuft of short to long, subapical spines............................................................ Kalimorpha Nielson

- Aedeagus more or less evenly tubular throughout, spines absent or variable in length and number, when present single or om row of short spines....................................................................... Jikradia Nielson

23. Pygofer in lateral view with or without caudoventral process, if present never with spines or teeth..................... 25

- Pygofer in lateral view with long broad, caudoventral process, process toothed or with spines in apical half..................................................................................................... Crepluvia Nielson

24. Subgenital plate elongate, sometimes broad medially; aedeagus with setae, spines or teeth never in row from base to near apex................................................................................................... 26

- Subgenital plate subglobular in apical half; aedeagus with row of setae from base to near apex........... Genatra Nielson

25. Aedeagus with setae and or spines on shaft................................................................ 31

- Aedeagus glabrous, toothed or flanged on shaft............................................................. 27

26. Aedeagus compressed laterally.......................................................................... 28

- Aedeagus not compressed laterally...................................................................... 29

27. Pygofer in lateral view with 2 caudal processes; aedeagus with toothed, medial flange.................. Jalorpa Nielson

- Pygofer in lateral view with single caudal process; aedeagus glabrous or toothed subapically............ Hastalidia Nielson

28. Style toothed or glabrous; aedeagus always more or less tubular............................................... 30

- Style usually with apical flange; aedeagus often inflated medially................................. Peayanus Nielson

29. Aedeagus glabrous; style triangulate in distal half with single, subapical tooth......................... Biadorus Nielson

- Aedeagus toothed subapically; style elongate................................................. Amylidia Nielson

30. Aedeagus broad subbasally, gonopore mostly subbasal, often large with numerous adjacent long setae, shaft often with setae distad of gonopore.................................................................................... 32

- Aedeagus narrow subbasally, gonopore subbasal, medial or subapical, shaft setaceous, rarely adjacent to subapical gonopore. .................................................................................................. 33

31. Style very slender, glabrous................................................................ Sapingia Nielson

- Style ornate often with numerous setae or from one to several spines............................... Stalolidia Nielson

32. Subgenital plate with median, longitudinal, double row of fine setae; pygofer with pendulate caudodorsal process.............................................................................................. Larsenolidia Nielson

- Subgenital plate and pygofer not as above................................................................. 34

33. Aedeagus with several to numerous spines or short spicules on shaft........................................... 37

- Aedeagus glabrous or with 1–2 subapical spines........................................................... 35

34. Aedeagus usually with, sometimes without, ventral subapical spine; subgenital plate often very broad in basal half............................................................................................... Docalidia Nielson

- Aedeagus with 2 subapical spines; subgenital plate often long, narrow, constricted in basal half....................... 36

35. Style long, robust, enlarged apically......................................................... Barodecus Nielson

- Style long, very narrow, sharply attenuated apically............................................... Baluba Nielson

36. Style long, narrow; aedeagus with setae or short spicules on shaft not as tuft adjacent to gonopore..................... 38

- Style very broad, plate-like; aedeagus with subapical gonopore and adjacent tuft of short setae....... Perspinolidia Nielson

37. Aedeagus setaceous on shaft; segment X glabrous........................................................... 39

- Aedeagus spiculated on middle of shaft; segment X with prominent ventral spine.................... Articoelidia Nielson

38. Medium size species, length less than 9 mm ............................................................... 40

- Large robust species, length more than 10 mm ............................................... Paraterulia Nielson

39. Style setaceous, spined or with row of teeth............................................................... 41

- Style glabrous and or with prominent spine................................................................ 42

40. Style with spine or teeth; subgenital plate setaceous......................................................... 43

- Style setaceous in distal 1/3; subgenital plate glabrous..................................... Neodocalidia , gen. nov.

41. Pygofer with caudoventral process when present not fused; style with spine medially or apically..................... 44

- Pygofer with large fused caudoventral lobe; style with row of teeth medially....................... Corupiana Nielson

42. Aedeagus with row of short setae medially; style with medial spine................................ Noritonus Nielson

- Aedeagus with row of long setae medially; style with 2 apical spines.............................. Plapigella Nielson

43. Subgenital plate distinctly constricted in subbasal third...................................................... 45

- Subgenital plate not as above........................................................................... 47

44. Aedeagus with several very long spines directed basally...................................................... 46

- Aedeagus with several short spines directed laterally or distally..................................... Jawigia Nielson

45. Subgenital plate elongate bulbous distally; aedeagus in lateral view narrowed in apical ¼.............. Onblavia Nielson

- Subgenital plate elongate-subtriangulate distally; adedeagus in lateral view very broad in apical 1/5...... Licontinia Nielson

46. Subgenital plate short, very broad....................................................... Marcapatiana Nielson

- Subgenital plate elongate............................................................................. 48

47. Aedeagus with long setae on shaft, removed from small gonopore.............................................. 49

- Aedeagus with very short setae adjacent to large gonopore..................................... Scapidonus Nielson

48. Aedeagus with coiled setae medially, 1 very long distal seta...................................... Inoclapis Nielson

- Aedeagus with row of setae of equal length............................................... Brevicapitorus Nielson

49. Aedeagus in lateral view with each appendage long, broad; dorsal appendage about ½ as long as ventral paraphysis, traversed by gonoduct.......................................................................... Korsigianus Nielson

- Aedeagus in lateral view with each appendage equal in length and configuration, dorsal appendage traversed by gonoduct....................................................................................... Spanigorlus Nielson

Figures

FIGURES 1–8. Freytagolidia ligula sp. nov. (1) pygofer and segment X, lateral view; (2) right subgenital plate, ventral view; (3) right style, dorsal view; (4) right style, lateral view; (5) aedeagus and dorsal connective, lateral view; (6) aedeagus and dorsal connective, ventral view; (7) connective, dorsal view; (8) dorsal connective, dorsal view.