Prototilla Schuster, 1949

Torréns, Javier, Fidalgo, Patricio, Roig-Alsina, Arturo & Brothers, Denis J., 2012, New species of Prototilla Schuster, 1949 from Argentina and diagnoses of the genus based on male and female (Hymenoptera: Bradynobaenidae: Typhoctinae: Eotillini), Zootaxa 3427, pp. 17-32 : 19

Prototilla Schuster, 1949

Prototilla Schuster 1949: 69 –70. Additional citations: Mickel 1968 (mention); Brothers 1970, 1974 (mention); Martinez & Fritz 1974 (mention); Fernandez 2000, 2001, 2002 (list); Brothers 2006 (key); Torréns & Roig-Alsina 2009 (mention).

Type species: Prototilla anomala Schuster, 1949 by original designation and monotypy.

Diagnosis. Male. Head with ocelli. Antennal scape smooth, without ventral longitudinal carina. Horizontal face of prosternum strongly projecting anteriorly, with a small semicircular projection posteriorly. Mid coxae moderately separated by a distance equal to half maximum width of femur. Fore wing with three submarginal cells ( Figs 6, 7, 10 ). Hind wing with vein 1RS proclinate, median cell apically produced, extending beyond origin of distal hamuli-group; median cell with three veins arising from it (RS, M and Cu) ( Fig. 32 ; see Fig. 9 in Schuster 1949). Tarsal claws each with ventral tooth (see Fig. 13 in Schuster 1949). T7 with surface convex (Figs 2, 5, 12). Genitalia with basal ring anteroventrally acutely emarginate ( Fig. 16 ).

Female. Head without ocelli. Pronotum rectangular with humeral tubercle pronounced and well defined ( Fig. 22 ). Mesopleural carina extending from anterior margin to posterior margin of sclerite ( Figs 25, 27 ). Mid coxae widely separated by a distance of 1.3 × maximum width of mid femur. Tarsal claws smooth, without teeth. S6 with latero-posterior margin smooth, without denticles.

Figures

FIGURES 6 – 10. Prototilla spp. 6, P. anomala, 3 (Santa Teresita, La Rioja), fore wing (lateral). 7 – 9, P. intercalata sp. nov., holotype 3 (Los Tigres, Santiago del Estero): 7, fore wing (lateral); 8, habitus (dorsal); 9, metasoma (lateral). 10, P. n i g r a sp. nov., 3 (Santa Teresita, La Rioja), fore wing (lateral).

FIGURES 29 – 35. Prototilla spp. 29 – 32, P. anomala, holotype 3 (Mendoza): 29, habitus (lateral); 30, habitus (dorsal); 31, wings and labels; 32, fore and hind wings. 33 – 35, P. typhoctoides, holotype Ƥ (Guanaco Muerto, Córdoba): 33, habitus (dorsal); 34, habitus (lateral); 35, labels.

FIGURES 11 – 16. Prototilla nigra sp. nov., holotype 3 (Lavalle, Mendoza): 11, habitus (lateral); 12, metasoma (dorsal); 13, habitus (dorsal); 14, head (anterior); 15, antenna (lateral); 16, genitalia (ventral; genital foramen projection indicated).

FIGURES 17 – 22. Prototilla spp. 17, 18, P. telteca sp. nov., paratype Ƥ (Lavalle, Mendoza): 17, habitus (lateral); 18, habitus (dorsal). 19, 20, P. striata sp. nov., paratype Ƥ (Lavalle, Mendoza): 19, habitus (lateral); 20, habitus (dorsal). 21, 22, P. typhoctoides, paratype Ƥ (Guanaco Muerto, Córdoba): 21, habitus (lateral); 22, habitus (dorsal; anterolateral angles of pronotum indicated).

FIGURES 23 – 28. Prototilla spp. 23, 24, P. t e l t e c a sp. nov., paratype Ƥ (Lavalle, Mendoza): 23, mesosoma (lateral); 24, head (anterior). 25, 26, P. striata sp. nov., paratype Ƥ (Lavalle, Mendoza): 25, mesosoma (lateral; longitudinal carina of mesopleuron indicated); 26, head (anterior). 27, 28, P. typhoctoides, paratype Ƥ (Guanaco Muerto, Córdoba): 27, mesosoma (lateral; longitudinal carina of mesopleuron indicated); 28, head (anterior).