Prototilla anomala Schuster, 1949

Torréns, Javier, Fidalgo, Patricio, Roig-Alsina, Arturo & Brothers, Denis J., 2012, New species of Prototilla Schuster, 1949 from Argentina and diagnoses of the genus based on male and female (Hymenoptera: Bradynobaenidae: Typhoctinae: Eotillini), Zootaxa 3427, pp. 17-32 : 22

Prototilla anomala Schuster, 1949

(Figs 1–3, 6, 29–32)

Prototilla anomala Schuster, 1949: 71 –75.

Diagnosis. Male. First and second segments of metasoma reddish, the rest black (Figs 1, 3, 29, 30). T1 and T2 with plumose setae across their surfaces, scale-like setae only on their posterior margins (Fig. 2). T1 spiracle located on a small protuberance. Fore wing moderately setose ( Figs 6 , 32 ). Female. Unknown.

Description and characterization provided by Schuster (1949).

Variation. T 3 may be entirely reddish like T1 and T2 (and then T 4 may be reddish basally), or reddish only basally, rather than entirely black.

Discussion. Schuster based his description on a specimen deposited in the Paris Museum (André Collection) from “ Argentina (no further data)”. In fact, the holotype bears a second label “Mendoza” ( Fig. 31 ), which narrows down the type locality. The additional specimens examined here extend the known distribution several hundred kilometers north-north-east of the type locality.

While the detailed description by Schuster (1949) is correct, a few additional characters useful to differentiate this species from others are given in the diagnosis above. Although Schuster indicated the length as “ 5–6 mm ”, which would indicate that he had more than one specimen, he stated (1949: 67, footnote) that Prototilla was represented by “the unique type ” and listed only the holotype, so the length was an estimate, perhaps because the metasomal segments are somewhat telescopically extended ( Figs 29, 30 ).

Distribution. Argentina: La Rioja, Mendoza.

Material examined. Holotype 3: ARGENTINA: Mendoza (no further data) ( MNHN). Other material, 6 3: La Rioja, Santa Teresita [28°35'57"S 66°33'32"W], 28/IX/2006, J. Torréns, G. Fidalgo, P. Fidalgo, MT (13, IFML); same location, 28/X/2006, G. y P. Fidalgo y J. Torréns, MT (13, MACN); same location, III/2005, MT, P. Fidalgo, G. Fidalgo y J. Torréns (13, MACN); 5 km N Carrizal, 490 m., malaise, 28°57.16'S 66°37.75'W, 2/ 16.ix.1999, ME Irwin, FD Parker (13, DJBC; 13, PAGL); Sierra de Argañaraz, 28 km N Chepes, Higuerita, malaise, 31°11.35'S 66°41.39'W, 2/ 16.ix. 1999, 930 m, ME Irwin, FD Parker (13, PAGL).

Figures

FIGURES 6 – 10. Prototilla spp. 6, P. anomala, 3 (Santa Teresita, La Rioja), fore wing (lateral). 7 – 9, P. intercalata sp. nov., holotype 3 (Los Tigres, Santiago del Estero): 7, fore wing (lateral); 8, habitus (dorsal); 9, metasoma (lateral). 10, P. n i g r a sp. nov., 3 (Santa Teresita, La Rioja), fore wing (lateral).

FIGURES 29 – 35. Prototilla spp. 29 – 32, P. anomala, holotype 3 (Mendoza): 29, habitus (lateral); 30, habitus (dorsal); 31, wings and labels; 32, fore and hind wings. 33 – 35, P. typhoctoides, holotype Ƥ (Guanaco Muerto, Córdoba): 33, habitus (dorsal); 34, habitus (lateral); 35, labels.