Eucoryphus Mik

Pusch, Martina H. E., Stark, Andreas & Pollet, Marc, 2020, Description of a new Eucoryphus species from the island of Corsica, France (Diptera: Dolichopodidae, Hydrophorinae), Zootaxa 4816 (4), pp. 527-540 : 529-530

Genus Eucoryphus Mik

Eucoryphus Mik, 1869: 19 . Type species: Eucoryphus brunneri Mik, 1869 (original designation).

Diagnosis. Medium-sized dolichopodids with body length of 2.5–4 mm. Frons broad, strongly widening above, antenna with pale yellowish basic colour, inserted in middle of the head, strongly modified in male: scape small, round (covered by pedicel in Eucoryphus piscariviverus sp. nov.), pedicel in E. coeruleus and E. brunneri reniform, extremely hypertrophied and barge- or bean-shaped in E. piscariviverus sp. nov.; postpedicel deeply bifurcate ( E. coeruleus , E. brunneri ) or more compact, flattened in lateral view, similar in shape as pedicel although much smaller ( E. piscariviverus sp. nov.); stylus simple ( E. coeruleus , E. brunneri ) or with a small cup-shaped basal part and bowed apical part ( E. piscariviverus sp. nov.). Eyes hairy, a pair of strong outer vertical setae, directed forward and inward. Cervix prolonged with a large triangular second cervical sclerite. Mesonotum without acrostichals, with six pairs of dorsocentral setae. Legs long and slender, sparsely bristled, in male with MSSC on tarsus of leg I; coxa III with a single outer seta. Abdomen with tergites 1–5 pubescent; in male sternite 1 with a soft outpouching, forming a wall- or edge-like structure; sternite 4 with dark and heavily sclerotized, V-shaped structure, appressed to the ventral side of the abdomen and bearing small stiff setae; tergites 4–5 prolonged laterally, tergite 6 hidden below tergite 5. Hypopygium of moderate size, compact and rather closely set to the abdominal segments. Cerci oval to elongate oval, with strong upcurved marginal setae. Autenna of female, as compared with that of male, rather simple, postpedicel more or less reniform. Six abdominal segments visible, tergite 10 bearing acanthophorites.

Remarks. The genus Eucoryphus is currently assigned to the subfamily Hydrophorinae within the family Dolichopodidae s. str. ( Sinclair & Cumming 2006). As interpreted by Negrobov (1981), Eucoryphus is part of the tribe Hydrophorini . Only Ulrich (1981) excluded Eucoryphus from the Hydrophorinae and classified it (along with Coracocephalus Mik, 1892 ) in the Sympycninae . This proposal, however, was not accepted by subsequent authors (e.g., Negrobov 1981; Yang et al. 2006; Germann et al. 2011). The newly species described below is assigned to Eucoryphus because it shares a unique combination of character states with the two described species: pedicel hypertrophied, postpedicel with long appendages (male) (synapomorphy), acrostichal setae lacking, six pairs of dorsocentral setae, tarsus of leg I with MSSC in male, and hind coxa with a single outer seta.