Holandriana holandri

Strong, Ellen E., 2011, More than a gut feeling: utility of midgut anatomy in phylogeny of the Cerithioidea (Mollusca: Caenogastropoda), Zoological Journal of the Linnean Society 162 (3), pp. 585-630 : 604

HOLANDRIANA HOLANDRI (PFEIFFER, 1828)

Material examined

Albania: Skoder Lake ( ZMB 106.145 ) .

Description

Marginal fold S-shaped ( Fig. 12A ), terminating at posterior tip of sorting area; oesophageal aperture at tip of marginal fold recurved segment; sorting area rectangular, with slightly curving, pointed posterior tip; left margin of sorting area straight; accessory marginal fold curving around posterior margin of sorting area, indistinct anteriorly due to poor preservation, intersecting short, slightly curving fold posteriorly at left; sorting area pad present at left posterior end of sorting area; crescentic pads lacking; glandular pad large, flaring posteriorly with broadly rounded posterior end, extending far posterior to gastric shield; single digestive gland duct opening at left of glandular pad; crescentic ridge long, extending from digestive gland duct, curving around posterior end of glandular pad, with distal tip curving deeply into caecum, forming shallow quarter loop under pad; caecum extending to midline of pad; single caecal fold present at right of glandular pad, behind caecum; accessory pads lacking; U-shaped fold present below style sac aperture; typhlosoles unfused.

Figures

Figure 12. Midgut anatomy of Melanopsidae. A, Holandriana holandri (ZMB 106.145). B, Melanopsis praemorsa (ZMB 106.066). Scale bars: 1.0 mm; refer to the text for a list of anatomical abbreviations.