Slaterocoris robustus (Uhler)

Schwartz, Michael D., 2011, Revision And Phylogenetic Analysis Of The North American Genus Slaterocoris Wagner With New Synonymy, The Description Of Five New Species And A New Genus From Mexico, And A Review Of The Genus Scalponotatus Kelton (Heteroptera: Miridae: Orthotylinae), Bulletin of the American Museum of Natural History 2011 (354), pp. 1-290 : 268-273

publication ID	https://doi.org/ 10.1206/354.1
DOI	https://doi.org/10.5281/zenodo.6864875
persistent identifier	https://treatment.plazi.org/id/0395E50B-FE86-FEFA-DFD6-FB79FE6EA72C
treatment provided by	Tatiana
scientific name	Slaterocoris robustus (Uhler)
status

Treatment

Slaterocoris robustus (Uhler) View in CoL

Figures 12 View Fig , 71–73 View Fig View Fig View Fig ; plates 3 View Plate 3 , 4M; map 13 View Map 13

Stiphrosoma robusta Uhler, 1895: 45 (orig. desc.).

Strongylocoris robustus: Van Duzee View in CoL : 1921: 136 (disc., in part).

Strongylocoris uniformis Van Duzee, 1925: 396 (orig. desc.); Kelton, 1968: 1132 (n. syn.).

Slaterocoris robustus: Kelton, 1968: 1132 View in CoL , fig. 19 (disc.); Knight, 1970: 238, fig. 1 (disc.); Henry and Wheeler, 1988: 451 (catalog); Schuh, 1995: 199 (catalog).

Slaterocoris nevadensis Knight, 1970: 239 View in CoL , fig. 20 (disc.); Henry and Wheeler, 1988: 450 (catalog); Schuh, 1995: 198 (catalog). NEW SYNONYMY.

DIAGNOSIS: Recognized by the uniform black coloration (pl. 3), except rarely for the pale apices of the femora (fig. 12); the sparse, or moderately sparse, long, erect, somewhat thickened, black vestiture (pl. 4M, fig. 71B); the relatively wide vertex (fig. 71A); and the male genitalia with the dorsal lobe of the endosomal spicule bifurcate and smooth marginally, the ventral lobe of the endosomal spicule smooth marginally (fig. 73A), and the relatively long, strongly (fig. 72B) or faintly bifurcate (fig. 72A) apex of the right paramere (fig. 72). The relatively stiff black vestiture (pl. 4M, fig. 71B) and smooth frons (fig. 73A) will easily distinguish S. robustus from all other congeners with completely black legs.

REDESCRIPTION: Male: Moderately small, obovate, length 3.35–5.13, width 1.65–2.18 (pl. 3). COLORATION: Shining black, including antenna and legs, except apex of femur rarely pale (fig. 12). VESTITURE: Sparse to moderately sparse, moderately long, erect, black bristlelike simple setae (pl. 4M, fig. 71B); antennal segments II to IV with erect, black setae. STRUCTURE: Body conformation in both sexes roughly similar; frons smooth (fig. 71A); labium reaching base of middle coxa; calli slightly produced, smooth, pronotal disc sparsely and shallowly punctate (fig. 71B); costal slightly arcuate. GENITALIA: Pygophore: Tergal process short with right side point, entire, directed toward midline. Phallotheca: Left lateral surface slightly compressed (fig. 73B). Endosomal spicule: Apical region of ventral lobe with uniform thickness throughout, distal to recurved angle smooth, long; dorsal lobe bifid with narrow lobes (fig. 73A). Right paramere: Small, apical region narrowed, medial region with prominent solitary spine (fig. 72A, arrow), basal lobe rounded with single (fig. 72A) or bifurcate (fig. 72B) spine, distal to basal lobe (fig. 72). Left paramere distal portion shorter than basal portion; ventral surface of equal diameter throughout (fig. 73C).

Female: Moderately small, ovate; length 3.40–4.35, width 1.70–2.40 (pl. 3). STRUCTURE: Costal margin strongly curved. GENITALIA: First gonapophyses: Left overlapping right, in ventral view, left greater than right at overlap; right lateral surface small. Left first gonapophyses: Dorsal surface concave, apex wide. Vestibulum, anteroventral margin of anterior wall: Sclerotized medially and sclerotized to lateral margins of bursa copulatrix. Ventral labiate plate: Strongly produced moderately, base wide, triangular. Second gonapophyses: Anterior medial surface slightly convex. Interramal sclerite: Dorsomedial region flat, not produced anteriorly, posteromedial portion with tumid process abutting ovipositor bulb, lateral ventromedial regions overlapping ventromedial plate. Interramal lobe: Dorsal or basal insertion with membranous attachment; dorsomedial margin not produced as lobe; ventral projection long with concave anterior surface.

HOSTS: The breeding host for S. robustus is Artemisia tridentata (Asteraceae) , including A. tridentata ssp. parishii , in California, and the widely distributed nominal subspecies. Many specimens were also collected from unidentified species of Artemisia throughout the distributional range of S. robustus . Fewer specimens were taken on five other Asteraceae : Achillea millefolium , Artemisia cana , A. frigida , Chrysothamnus viscidiflorus , and unknown species of Chrysothamnus . The limited number of non-Asteraceae records are considered sitting records and not indicative of breeding hosts.

In central Oregon, S. robustus (identified as S. nevadensis ) was collected along with three phyline plant bugs from Artemisia tridentata as part of a study of the effects of secondary plant chemistry and bird predation on the sagebrush arthropods (Wiens et al., 1991).

DISTRIBUTION: Western North America from the Okanagan Valley of southern British Columbia southeast to the central Rocky Mountains of Colorado and southwest to southern California ( map 13 View Map 13 ). Although widely cited (e.g., Kelton, 1968; Knight, 1970; Henry and Wheeler, 1988), I have not located specimens of S. robustus from Texas.

COLLECTION SUMMARY: 985 from 226 collection events. Earliest collections in May are from California and the Columbia Basin of Washington. Most collection events are from June and July with August dates predominating at higher latitude and elevation.

DISCUSSION: Kelton (1968) designated a specimen with the labels ‘‘ Stiphrosoma robusta Uhl., Colo. 191; P.R. Uhler Collection; No. 69538’’ as the lectotype of Stiphrosoma robusta (USNM) . This specimen could not be found in the holdings of the CNC or the running and type collections of the USNM. Uhler (1895) described Stiphrosoma robusta based on two females from Colorado. There are several specimens in the USNM with a least these two labels: ‘‘Colo [printed] ### (#) [handwritten]’’ and ‘‘PR Uhler Collection’’ (# 191 [♀], # 1329 [8], # 1332 [8], # 1412 [2♀], # 1581 [98, 13♀], # 1686 [8, ♀], # 2010 [♀], # 2012 [8, 4♀], # 2158 [48, 3♀], # 2193 [28, 3♀]). The # 191 female has three additional labels: ‘‘ Stiphrosoma robusta Uhler’’ in Uhler’s handwriting; ‘‘ Stiphrosoma robusta Uhler’’ [handwritten] and ‘‘Det UHLER’’ [printed]; and a red label. This is probably the specimen designated by Kelton (1968), although in the note book of Gillette and Baker collection codes held in the USNM, # 191 references a specimen from ‘‘M.A.C.’’ [Michigan Agriculture College?], ‘‘breeding cage,’’ ‘‘5–20–90, ‘‘C.F. B[aker].’’ None of the specimens numbers listed above exactly matches the original description of ‘‘Steamboat Springs, July 26th (Gillette).’’ As # 191 agrees with the original description and concept of robusta , I have applied two labels to it in order to stabilize the intent of Uhler and clarify the ambiguous lectotype designation of Kelton: AMNH_PBI 00134033 and LECTOTYPE ♀ Stiphrosoma robusta Uhler, 1895 , desig. by M.D. Schwartz, 2010 [red label].

Knight (1970) described the nominal species S. nevadensis from two males collected near Las Vegas, Nevada. In his key S. nevadensis and S. robustus are separated by a comparison of antennal segment II length to vertex and eye width and male body length. The segment is subequal to the width of the vertex and an eye and the body is 4.20 in S. robustus ; in S. nevadensis the segment is not equal to the combined width and the length is 3.20. The form of the right paramere was also presumed to be diagnostic for the two nominal species. Having examined large numbers of specimens, including the types and associated material, from a variety of localities and hosts, I cannot find constant differences in body length, ratio of antennal segment II to head, or genitalic structure to support the distinction. My observations show that S. robustus has great size variation. The male body length ranges from 3.35 to 5.13 and the ratio of the head width to length of antennal segment II is from 1.12 to 1.63. Even though Knight (1970) illustrated the right paramere of the two nominal species, my comparisons suggest that these renderings do not document the variation that actually exists (cf. fig. 72A–F). There are no discrete differences in body size, head, vertex, and pronotal width, antennal segment II length, and structure of the right paramere that are correlated or tied to a particular geographical region. Short body length is found in specimens from southern Nevada as well as central Oregon (Silver and Summer lakes). Generally smaller specimens occur at lower elevations (approx. 1300 m) in and around the Great Basin whereas larger specimens are found at higher elevation (approx. 2700 m) and higher latitude (approx. 446 N). The long, moderately sparse, black vestiture of the dorsum, black body and appendages, relatively short antennal segment II, and the structure of the endosomal spicule as in figure 73A are unique and sufficient to diagnose this species. I therefore treat S. nevadensis as a junior synonym of S. robustus .

There is variation in the shape of the right paramere, especially in the number of apical spines and apical structure of the spine adjacent to the basal lobe. The apical region can be entire (fig. 72E) or with bifurcate spines of variable size and separation (fig. 72A–D, E). Neither Kelton (1968) or Knight (1970) documented the two small apical spines as shown in figure 72A, E, from Gould and Steamboat Springs, Colorado, respectively. Two large apical spines, as seen in figure 72B for a Nevada specimen, were also found in some specimens from British Columbia . The basal spine can also be entire (fig. 72A, D–F) or bifurcate (fig. 72B, C). Nonetheless, the structure of the endosomal spicule is consistent in all specimens examined from across the range of robustus ; all are nearly identical to figure 73A. I do note that the curvature of the long ventral lobe can change depending on the storage medium.

Based on the cladistic analysis, S. robustus was not placed in a species group, but either as the sister to the flavipes group (fig. 1B, D) or as the sister to a large number of species ( fig. 2 View Fig ), sometimes including S. ambrosiae (fig. 1A). Different sets of character pairs unambiguously support each of these arrangements of species. Black legs (5-2, homoplasious; 6-1, nonhomoplasious) partially define the node containing S. robustus and the argenteus , hirtus , and punctatus groups ( fig. 2 View Fig ). Ambiguous support for this arrangement is provided by vestiture coloration (9-0) and the structure of the anterior portion of the right paramere (40-1).

The smooth distal portion of the ventral lobe of endosomal spicule (34-1) and the left first gonapophysis overlapping the right (51- 0) gives nonhomoplasious support to the sister relationship of S. robustus and the flavipes group (fig. 1B). Additionally, vestiture coloration (9-0) and the structure of the anterior portion of the right paramere (40-2) provide ambiguous support to this arrangement. An arrangement (fig. 1A) where S. robustus is the sister to S. ambrosiae and the apache , argenteus , hirtus , and punctatus groups receives homoplasious support from the structure of the dorsal lobe of the endosomal spicule (36-3) and the ventral labiate plate (59-1). Depending on the optimization, ambiguous support for this arrangement is provided by two to five additional characters.

LECTOTYPE: USA: Colorado: Unknown co.: ‘‘ Colo 191,’’ unknown, 1♀ ( AMNH _ PBI 00134033 About AMNH ) ( USNM).

HOLOTYPES: Nevada: Clark Co.: Moapa [36.675556 N 114.620556 W], 25 May 1963, R.C. Bechtel, 18 ( AMNH _ PBI 00122256 About AMNH ) ( USNM) ( S. nevadensis : junior synonym) GoogleMaps ; Utah: Wasatch Co.: Heber [City] [40.506946 N 111.41256 W], 05 Jul 1922, E.P. Van Duzee 18, not examined ( CAS) ( S. uniformis : junior synonym) GoogleMaps .

OTHER SPECIMENS EXAMINED: CANADA: British Columbia : 4 mi E of Cawston, 49.191756 N 119.72376 W, 09 Jul 1959, L.A. Kelton, 18 (00111004) (CNC). Mount Kobau, 11 km NW of Osoyoos, 15 km SE of Summit, 49.101976 N 119.614096 W, 964 m, 30 Aug 1993, M.D. Schwartz, Artemisia tridentata , 48 (00110982, 00111005–00111006, 00115512) 3♀ (00111007–00111009) (CNC). Osoyoos [on Rt 3], 49.008456 N 119.409876 W, 560 m, 08 Jul 1991 – 3 Jul 1991, unknown, 18 (RBCM). USA: Arizona: Coconino Co.: Jacob Lake, 34.321116 N 110.846 W, 05 Jun 1969, W.F. Chamberlain, 1♀ (00118262) (TAMU). California: Alpine Co.: Ebbetts Pass, Pacific Crest National Scenic Trail, Toiyabe National Forest on Rt 4, 38.544446 N 119.810836 W, 2661 m, 05 Jul 1994, M.D. Schwartz, Artemisia sp. , 2♀ (00111012, 00111013) (CNC). Monitor Pass, 38.675566 N 119.619446 W, 06 Aug 1962, D.R. Smith, 18 (00118068) (ORSU). Monitor Pass, just W of, on Rt 89, 38.665336 N 119.641166 W, 2550 m, 27 Jul 1999, M.D. Schwartz, Chrysothamnus viscidiflorus , 28 (00111010–00111011) (CNC). Kern Co.: E of Caliente on Walker Basin Road, 35.387746 N 118.4256 W, 1220 m, 22 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Artemisia tridentata ssp. parishii , det. A. Sanders UCR 140648, 48 (00108002–00108005) (AM), Artemisia tridentata ssp. parishii , det. A. Sanders UCR 140648, 28 (00107999–00108000), 1♀ (00108001) (AMNH). S of Tehachapi on Water Canyon Rd, 35.095636 N 118.49096 W, 1445 m, 21 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Artemisia tridentata ssp. parishii , det. A. Sanders UCR 140648 [PBI CAL04-H5], 18 (00137233) (AMNH). nr Walker Pass, 35.66256 N 118.025836 W, 1524 m, 30 May 1981, J.T. Polhemus, 1♀ (00116754) (JTP). Lassen Co.: Blue Lake, 41.148896 N 120.289726 W, 19 Jun 1947 – 20 Jun 1947, R.L. Usinger, 18 (00119265) (UCB). Martin Springs, T31N R9E Sec 14, 40.551816 N 120.926316 W, 1830 m, 23 Jul 1922, J.O. Martin, 1♀ (00242622) (CAS). Modoc Co.: 2.5 mi S of Rt 139 toward Lookout, 41.171916 N 121.154176 W, 1440 m, 06 Jul 1979, R.T. and Joe Schuh, Artemisia tridentata , 28 (00107862, 00108014), 7♀ (00108015–00108021), Lupinus sp. (Fabaceae) , 18 (00108013) (AMNH). 24.7 mi NW of Canby, 41.696386 N 121.207326 W, 1375 m, 01 Jul 1979, R.T. Schuh and B.M. Massie, 1♀ (00108032) (AMNH). W of Cedarville, Cedar Pass, 41.56256 N 120.267786 W, 1850 m, 01 Jul 1979, R.T. Schuh and B.M. Massie, Artemisia tridentata , 38 (00108008– 00108010), 2♀ (00108011–00108012) (AMNH). Mono Co.: Bridgeport, 38.255836 N 119.230286 W, 28 Jul 1963, W.F. Chamberlain, 1♀ (00118231) (TAMU). Coleville, 38.566396 N 119.506116 W, 1585 m, 10 Jun 1966, W. Gagne, 38 (00119230–00119232), 2♀ (00119233–00119234) (UCB); 06 Jul 1966, C.W. O’Brien, 1♀ (00119267) (UCB). Conway Pass, 38.088066 N 119.180836 W, 18 Jun 1959, Frank M. Beer, 1♀ (00118072) (ORSU). Devils Gate Summit, 38.414176 N 119.169726 W, 2292 m, 06 Jul 1966, L. and C.W. O’Brien, 38 (00119225, 00119235–00119236), 3♀ (00119237–00119239) (UCB). Mono Lake, N edge at Rt 395, 38.073156 N 119.167456 W, 2187 m, 11 Jul 1980, R.T. Schuh and G.M. Stonedahl, Artemisia tridentata , 3♀ (00108027– 00108029) (AMNH). Toms Place, 37.561396 N 118.680286 W, 09 May 1966, unknown, 28 (00119240–00119241), 5♀ (00119242–00119246) (UCB). Virginia Lakes, 38.050126 N 119.256546 W, 2972 m, 15 Aug 1962, H.B. Leech, 18 (00242620) (CAS). Nevada Co.: 7 mi N of Truckee, Sagehen Creek, 39.438546 N 120.193376 W, 29 Jul 1962, J.T. Doyen, 18 (00119264) (UCB). 17 mi N of Truckee, 39.574486 N 120.182226 W, 21 Jul 1970, H.D. Hertert, 18 (00122330) (UID). Donner Lake, 39.32256 N 120.265286 W, 28 Jul 1932, R.L. Usinger, Artemisia tridentata , 18 (00119247), 10♀ (00119248–00119257) (UCB). Truckee, Tahoe National Forest, 39.328066 N 120.182226 W, 29 Aug 1967, L.A. Kelton, Artemisia sp. , 9♀ (00111014–00111022) (CNC). Riverside Co.: Hemet Reservoir, San Jacinto Mountains, 33.74756 N 116.971116 W, 22 May 1940, C.D. Michener, Artemisia tridentata , 28 (00119224, 00119260), 1♀ (00119261) (UCB). Herkey Creek, San Jacinto Mountains, 33.669176 N 116.680286 W, 04 Jun 1940, unknown, Adenostoma sp. (Rosaceae) , 1♀ (00242621) (CAS); 04 Jun 1940, C.D. Michener, 1♀ (00119262) (UCB); 04 Jun 1940, R.L. Usinger, Arctostaphylos sp. (Ericaceae) , 1♀ (00119263) (UCB). San Bernardino Co.: Cajon Pass, 34.326456 N 117.428696 W, 1177 m, 06 Jun 1935, P.W. Oman, 1♀ (00134528) (USNM). E of Wrightwood on Rt 2, 2 km W of Rt 138, Los Angeles National Forest, 34.378246 N 117.61176 W, 1556 m, 18 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, 18 (00091705) (AM). Mojave River Forks, 10 km SE of Hesperia, 34.349276 N 117.240646 W, 901 m, 14 May 1985, M. Clement, 18 (00119803) (UCR); 06 May 1986, Anja Lachmann (for Ent. 109), 18 (00119796) (UCR). Oro Grande Wash, 11 mi S of Adelanto, 34.421336 N 117.403916 W, 26 May 1941, Timberlake, Artemisia tridentata , 28 (00119801–00119802) (UCR). Just E of Wrightwood on Rt 2, 34.357746 N 117.61056 W, 1700 m, 18 May 2004, Schuh, Cassis, Schwartz, Weirauch, Wyniger, Forero, Artemisia tridentata ssp. parishii , det. A. Sanders UCR 140648, 18 (00108006), 1♀ (00108007) (AM), Artemisia tridentata ssp. parishii , det. Field ID, 198 (00322776, 00322777, 00322780– 00322796), 5♀ (00322773–00322775, 00322778, 00322779) (AMNH). San Diego Co.: Buckmans Springs, 32.771526 N 116.4926 W, 1021 m, 06 Jun 1928, C.C. Searl, Arctostaphylos sp. (Ericaceae) , 1♀ (00244260) (SDNH); 19 Jun 1951, C.F. Harbison, 1♀ (00244259) (SDNH). Morena Dam, 32.780836 N 117.206946 W, 26 May 1929, C.C. Searl, 38 (00244245–00244247), 4♀ (00244248–00244251) (SDNH). Pine Valley, 32.835836 N 116.533616 W, 1190 m, 13 Jun 1927, C.C. Searl, 28 (00244252–00244253), 4♀ (00244254–00244257) (SDNH); 04 Jun 1928, C.C. Searl, 1♀ (00244261) (SDNH); 11 Jun 1953, C.F. Harbison, 1♀ (00244258) (SDNH). Warner Springs, 33.049696 N 117.042336 W, 29 May 1978, Brown and Faulkner, 78 (00244238–00244244) (SDNH). Sierra Co. : 5 mi S of Sierraville, 39.517246 N 120.366396 W, 18 Jun 1966, C.W. O’Brien, Artemisia tridentata , 18 (00119258), 1♀ (00119259) (UCB). Siskiyou Co.: 2 mi N of Tule Lake Refuge Headquarters, 41.938356 N 121.531946 W, 1219 m, 26 Jun 1979, G.M. Stonedahl, Artemisia tridentata , 1♀ (00108031) (AMNH). Grass Lake, 41.6356 N 122.190286 W, 1524 m, 04 Jul 1952, M.A. Cazier, W. Gertsch, and R. Schrammel, 18 (00108033) (AMNH). Lava Beds National Monument near Headquarters, 41.753336 N 121.505566 W, 1560 m, 26 Jun 1979, R.T. and Joe Schuh, Artemisia tridentata , 1♀ (00108030) (AMNH). Macdoel, 41.826946 N 122.004176 W, 01 Jul 1935, P.W. Oman, 18 (00134539) (USNM). Shovel Creek Meadow, Willow Creek Mountain, 41.971116 N 122.201416 W, 10 Aug 1962, W. Peters and J. Schuh, 18 (00118069), 2♀ (00118070–00118071) (ORSU). Tulare Co.: 21 mi W of Rt 395 toward Kennedy Meadows, 36.009446 N 118.503086 W, 1875 m, 01 Jul 1980, R.T. Schuh, Artemisia tridentata , 5♀ (00108022–00108026) (AMNH). Ventura Co.: 18 mi E of Camp Ozena, 34.412376 N 119.039576 W, 02 Jul 1965, E.M. Omi, 1♀ (00119266) (UCB). Colorado: Boulder Co.: Bald Mountain, Boulder, 39.994446 N 105.610286 W, 2134 m, 17 Jun 1961, J.E.R. Stainer, 1♀ (00111243) (CNC). Clear Creek Co.: Berthoud Pass, 39.79816 N 105.778066 W, 3455 m, 12 Jul 1949, J.R. White , 1♀ (00117810) (KU). Eagle Co.: 22 km N of Wolcott, Big Piney Creek, 39.900286 N 106.678066 W, 27 Jun 1989, D.A. Polhemus, Artemisia tridentata , 38 (00116703, 00116737– 00116738), 1♀ (00116739) (JTP). N of Eagle, 39.655286 N 106.828066 W, 24 Jun 1979, J.T. Polhemus, 3♀ (00116742–00116744) (JTP). Shrine Pass Road, nr Vail, 39.640286 N 106.373616 W, 12 Aug 1987, D.A. Rider, 1♀ (00242957) (DAR). S of Wolcott, 39.702786 N 106.678066 W, 22 Jun 1980, J.T. Polhemus, 28 (00116740–00116741) (JTP). State Bridge, nr Bond, 39.857786 N 106.649176 W, 2134 m, 24 Jun 1961 – 25 Jun 1961, J.R. Stainer, 168 (00110985, 00111220–00111233, 00111235), 2♀ (00111234, 00111236) (CNC), 18 (00121782) (UCB). Vail, 39.640286 N 106.373616 W, 2591 m, 23 Jun 1986, J.T. Polhemus, Artemisia tridentata , 48 (00116732–00116734, 00116736), 1♀ (00116735) (JTP). Water Wheel Ranch near Bond, 39.874446 N 106.686676 W, 24 Jun 1978, J.T. Polhemus, 18 (00116747) (JTP). Garfield Co.: Baxter Pass (Radio Tower), 14 mi SE of CO State Border, 39.581946 N 108.951946 W, 2499 m, 09 Jul 1981, M.D. Schwartz, Artemisia tridentata , 88 (00110984, 00111026, 00111263–00111268), 13♀ (00111030– 00111032, 00111269–00111278) (CNC). W Evacuation Creek, 11 mi SE of Colorado state line on Rt 45, 39.772866 N 109.048356 W, 2073 m, 09 Jul 1981, M.D. Schwartz, Artemisia tridentata , 3♀ (00111282–00111284) (CNC). Grand Co.: Granby, 40.086116 N 105.938896 W, 12 Jun 1963, H.F. Howden, 18 (00111172) (CNC). Grand Lake Entrance, Rocky Mountains National Park, 40.252226 N 105.82256 W, 18 Aug 1968, L.A. Kelton, Artemisia sp. , 88 (00111173–00111180), 39♀ (00111181– 00111219) (CNC). Tabernash, 40.010146 N 105.920716 W, 2613 m, 08 Aug 1973, D. Kopp and J. Aldrich, 18 (00243060), 1♀ (00243061) (UMRM). [Big] Muddy Pass, US Rt 40, 40.37866 N 106.5796 W, 2652 m, 12 Aug 1973, T.R. Yonke, 18 (00243059) (UMRM). Gunnison Co.: 28 mi E of Montrose on Rt 50, 38.477186 N 107.357016 W, 2621 m, 17 Jul 1986, R.T. Schuh, Artemisia tridentata , 28 (00107929–00107930) (AMNH). 35 mi E of Gunnison on Rt 50, 38.545836 N 106.277776 W, 2591 m, 17 Jul 1986, R.T. Schuh, Artemisia tridentata , 48 (00107870, 00107935–00107937), 3♀ (00107938–00107940) (AMNH). Almont, 38.664446 N 106.846116 W, 2743 m, 19 Jun 1925, A.W. Lindsey, 98 (00122149–00122157) (USNM). Red Creek and Gunnison River, 38.468086 N 107.240346 W, 2262 m, 29 Jun 1961, E. Riley, 18 (00134531) (USNM). West Elk Creek Junction, 38.715276 N 107.205886 W, 2256 m, 03 Jul 1961, E. Riley, 18 (00134518) (USNM). Jackson Co.: 4 mi N of Gould, 40.584216 N 106.026116 W, 05 Aug 1975, J.C. Schaffner, 2♀ (00116748–00116749) (JTP), 118 (00118242– 00118245, 00118282–00118288), 20♀ (00118246, 00118263–00118281), Artemisia cana , 118 (00118217, 00118228–00118230, 00245339– 00245345), Artemisia frigida , 1♀ 268 (00094016–00094041), 55♀ (00093716–00093717, 00093798–00093808, 00094047–00094087, 00245338) (TAMU). 7 mi NE of Rand, 40.525446 N 106.086696 W, 14 Aug 1969, J.C. Schaffner, 28 (00245348–00245349), 2♀ (00245350–00245351) (TAMU). Cowdrey, 40.859726 N 106.31256 W, 2438 m, 15 Aug 1968, L.A. Kelton, Chrysothamnus sp. , 18 (00111171) (CNC). Gould, 40.526396 N 106.026116 W, 2804 m, 12 Aug 1968 – 13 Aug 1968, L.A. Kelton, Artemisia sp. , 168 (00110983, 00110990, 00111133, 00111158– 00111170) (CNC). Muddy Pass, Routt National Forest, 40.376676 N 106.579176 W, 2652 m, 19 Aug 1968, L.A. Kelton, Artemisia sp. , 58 (00111124–00111128) (CNC); 21 Aug 1968, L.A. Kelton, Artemisia sp. , 118 (00111100–00111110), 13♀ (00111111–00111123) (CNC). North Park, 40.90006 N 106.308336 W, Jul 1926 – Aug 1926, F.C. Hottes, 1♀ (00134136) (USNM). Rand, 40.453566 N 106.179066 W, 2627 m, 26 Aug 1940, H.C. Severin, 1♀ (00134519) (USNM). nr Cowdrey, Colorado-Wyoming state line on Rt 125, 40.859726 N 106.31256 W, 04 Aug 1979, J.T. Polhemus, 18 (00116745), 1♀ (00116746) (JTP). Jefferson Co.: Deer Creek Canyon, 39.552656 N 105.120846 W, 1981 m, 11 Jul 1986, R.T. Schuh, J.T. Polhemus, Artemisia tridentata , 1♀ (00107941) (AMNH). Moffat Co.: Escalante Overlook on Harpers Corner Road, 8.1 miles N route 40, 40.326116 N 108.9456 W, 2350 m, 08 Jul 1982, M.D. Schwartz, Artemisia tridentata , 38 (00111279– 00111281) (CNC). Montezuma Co. : Mesa Verde National Park, 37.233336 N 108.479176 W, 2134 m, 01 Jun 1927, V.M. Tanner, 18 (00134511) (USNM). Montrose Co.: Black Canyon of Gunnison, 38.586 N 107.756 W, 18 Aug 1984, R.L. Blinn, Artemisia sp. , 1♀ (00242849) (NCSU). Black Canyon of the Gunnison Wilderness, 38.584446 N 107.756666 W, 15 Aug 1973, J.C. Schaffner, 18 (00118292), 3♀ (00118289–00118291) (TAMU). Routt Co.: Steamboat Springs, 40.4856 N 106.831116 W, 2134 m, 30 Jun 1931, R.H. Beamer, 48 (00117803–00117806) (KU); 15 Jul 1964, H.H. Knight, 18 (00111130), 1♀ (00111132) (CNC); 16 Jul 1964, H.H. Knight, 18 (00111129), 1♀ (00111131) (CNC), 18 (00118236), 2♀ (00118237–00118238) (TAMU). Steamboat Springs, 40.4856 N 106.831116 W, 2103 m, 11 Jul 1964, H.H. Knight, 38 (00134502–00134504) (USNM). Steamboat Springs, 40.4856 N 106.831116 W, 2042 m, 11 Jul 1964, H.H. Knight, 18 (00118241) (TAMU); 15 Jul 1964, H.H. Knight, 28 (00118239–00118240) (TAMU). Saguache Co.: 23 mi E of Gunnison on Rt 50, 38.41086 N 106.503026 W, 2469 m, 17 Jul 1986, R.T. Schuh, Artemisia tridentata , 18 (00107931), 3♀ (00107932–00107934) (AMNH). Summit Co.: Dillon, 39.630286 N 106.042786 W, 29 Aug 1972, L.A. Kelton, Artemisia sp. , 38 (00111237–00111239), 3♀ (00111240– 00111242) (CNC). Weld Co.: 5 mi N of Nunn, 40.774576 N 104.789636 W, 1629 m, 09 Aug 1973, T.R. Yonke, 1♀ (00243062) (UMRM). Idaho: Ada Co. : Regina, 43.39216 N 115.996 W, 1036 m, 18 Jun 1962, W.F. Chamberlain, 38 (00245335–00245337), 2♀ (00245346–00245347) (TAMU). Adams Co.: 7 mi E of New Meadows, 44.97136 N 116.139776 W, 02 Aug 1972, W.F. Barr, Artemisia tridentata , 38 (00122318–00122320), 1♀ (00122321) (UID). Bannock Co.: 3 mi E of McCammon, 42.650566 N 112.133256 W, 29 Jun 1966, W. Gagne and J. Haddock, 18 (00119273) (UCB). Bear Lake Co.: Cub River Canyon, Thompson Spring, 42.548826 N 111.293266 W, 2275 m, 05 Jul 1968, G.F. Knowlton and H.H. Knight, 18 (00134408) (USNM). Blaine Co.: 6 mi NW of Carey, Little Wood River, 43.378916 N 113.997586 W, 30 Jun 1966, W. Gagne and J. Haddock, 1♀ (00119272) (UCB). Sun Valley, 43.688756 N 114.317726 W, 2084 m, 18 Jul 1970, K.E. Gibson, 1♀ (00134137) (USNM). Boise Co.: Mores Mountain, 43.784316 N 116.086166 W, 2103 m, 09 Jul 2006, A.R. Gillogly, 48 (00094008 [cat TAMU-ENTO X0282229]– 00094011 [cat TAMU-ENTO X0283444]), 3♀ (00094012 [cat TAMU-ENTO X0282357]– 00094014 [cat TAMU-ENTO X0283962]) (TAMU). Butte Co.: 7 mi E of Howe, 43.783526 N 112.863456 W, 22 Jun 1966, W.F. Barr, 18 (00122329) (UID). Craters of the Moon National Monument, 43.429726 N 113.530566 W, 13 Jul 1965, D.S. Horning, Jr., Artemisia tridentata , 18 (00134532) (USNM). Craters of the Moon National Monument, 43.416676 N 113.516676 W, 23 Jul 1982, J. B Johnson, R.L. Gillespie , 18 (00122332), 1♀ (00122438) (UID). Camas Co.: 12 mi S of Fairfield, 43.209626 N 114.786996 W, 11 Jul 1978, unknown, 18 (00122327) (UID). Custer Co.: Bonanza, 44.370566 N 114.726946 W, 22 Jul 1965, W.F. Barr, 18 (00122331) (UID). Clayton, 44.258226 N 114.401246 W, 1658 m, 07 Jul 1928, C. Wakeland, 18 (00134517) (USNM). Elmore Co.: 10 mi S of Sunnyside, 43.139036 N 115.952786 W, 25 Jun 1964, W.F. Barr, Artemisia tridentata , 18 (00122326) (UID). 11 mi S of Pine, 43.324836 N 115.311116 W, 22 Jun 1977, W.F. Barr, Vernonia sp. (Scrophulariaceae) , 18 (00122328) (UID). 13 mi N of Mountain Home, 43.321376 N 115.690286 W, 18 Jun 1966, A.R. Gittins, Achillea millefolium , 18 (00122325) (UID). Nez Perce Co.: Beaver Canyon, 46.397646 N 116.80796 W, 317 m, 23 Jul, unknown, 1♀ (00134513) (USNM). Owyhee Co.: 5 mi N of Reynolds, 43.260766 N 116.710346 W, 1249 m, 07 Jul 1965, W.F. Barr, 18 (00122436) (UID). 6 mi N of Reynolds, 43.257976 N 116.710446 W, 1259 m, 07 Jul 1965, W.F. Barr, 1♀ (00122437) (UID). 6 mi SE of Grasmere, 42.315176 N 115.798466 W, 06 Jul 1965, R.L. Westcott, 18 (00122322), 2♀ (00122323–00122324) (UID). Grand View Road, 42.992266 N 116.097066 W, 718 m, 25 Jun 1964, W.F. Barr, 18 (00122431), 4♀ (00122432–00122435) (UID). Power Co.: American Falls, 42.786116 N 112.853616 W, 05 Aug 1972, L.A. Kelton, Artemisia sp. , 88 (00111088–00111095), 4♀ (00111096–00111099) (CNC). Teton Co.: Pine Creek Pass, 43.571676 N 111.214446 W, 06 Aug 1972, L.A. Kelton, Artemisia sp. , 118 (00111070–00111080), 7♀ (00111081–00111087) (CNC). Twin Falls Co.: Roseworth, 42.366946 N 114.921396 W, 16 Jul 1963, O.O. Fillmore, G.B. Hewitt, 1♀ (00122333) (UID). Montana: Carbon Co.: 11 mi S of Red Lodge, Parkside Campground, 45.062216 N 109.410246 W, 2210 m, 12 Aug 1986, M.D. Schwartz and G.M. Stonedahl, Artemisia tridentata , 28 (00107867, 00107919), 2♀ (00107921–00107922) (AMNH). Gallatin Co.: Beaver Creek, 44.856876 N 111.373566 W, 1920 m, 15 Aug 1913, S.J. Hunter, 18 (00117809) (KU). Elkhorn Ranch, Gallatin River, Upper Gallatin Canyon, 45.067866 N 111.187636 W, 2134 m, 01 Aug 1928 – 05 Aug 1928, J. McDunnough, 148 (00110986, 00111035–00111047), 1♀ (00111048) (CNC). Targhee Pass on Rt 191, 44.674726 N 111.2756 W, 2156 m, 10 Aug 1986, Schuh, Schwartz, and Stonedahl, Artemisia tridentata , 28 (00107920, 00107923), 3♀ (00107924– 00107926) (AMNH). Nevada: Carson City Co.: Carson City, 39.163896 N 119.766396 W, 1829 m, 25 Jun 1929, R.L. Usinger, 28 (00242591–00242592) (CAS); 26 Jun 1929, R.L. Usinger, 18 (00242593), 2♀ (00242594– 00242595) (CAS), 18 (00134538) (USNM). Clark Co.: Moapa, 36.675556 N 114.620556 W, 25 May 1963, R.C. Bechtel, paratype of Junior Synonym ( S. nevadensis ), 18 (00134017) (USNM). Elko Co.: 30 mi SE of I-80 on Hwy 229, 40.638826 N 115.266766 W, 1908 m, 19 Jul 1980, G.M. Stonedahl, Artemisia sp. , 2♀ (00108093–00108094) (AMNH). Ruby Mountains, Lomoille Canyon, 40.960566 N 114.949176 W, 1890 m, 16 Jul 1970, D.G. Denning, 2♀ (00108091– 00108092) (AMNH). Mineral Co.: 27 mi SW of Hawthorne on Rt 359, 1 mi NE of Anchorite Summit, 38.248716 N 118.975066 W, 2256 m, 02 Jul 1983, R.T. Schuh and M.D. Schwartz, Artemisia tridentata , 18 (00107944), Chrysothamnus viscidiflorus , 1♀ (00107943), Purshia glandulosa Curran (Rosaceae) , 18 (00107942) (AMNH). Nye Co.: 1 mi NE of Belmont on Rt 82, 38.606336 N 116.860246 W, 2280 m, 13 Jul 1980, R.T. Schuh and G.M. Stonedahl, Artemisia tridentata , 18 (00108095) (AMNH). 3.5 mi SE of Manhattan, Toiyabe National Forest, 38.503116 N 117.026786 W, 2134 m, 13 Jul 1980, R.T. Schuh and G.M. Stonedahl, Artemisia tridentata , 58 (00107866, 00108076–00108079), 11♀ (00108080–00108090), Lupinus sp. (Fabaceae) , 28 (00108074–00108075) (AMNH). Washoe Co.: 6.2 mi W of Junction of Rts 8A and 34A, 41.581996 N 119.684116 W, 1985 m, 02 Jul 1979, R.T. Schuh and B. Massie, Artemisia tridentata , 18 (00108096), 2♀ (00108097–00108098) (AMNH). 7 mi W of Vya on Rt 8A toward Cedarville, 41.592226 N 119.994756 W, 1800 m, 02 Jul 1979, R.T. Schuh and B.M. Massie, Artemisia tridentata , 318 (00108099–00108129), Chrysothamnus sp. , 18 (00107871) (AMNH). 17 mi NW of Gerlach, 40.825656 N 119.583796 W, 11 Jun 1970, unknown, 18 (00119268), 1♀ (00119269) (UCB). White Pine Co. : Wheeler Peak Scenic Drive, Humboldt National Forest, 38.986116 N 114.31256 W, 2591 m, 06 Aug 1982, M.D. Schwartz, Artemisia tridentata , 48 (00111027, 00111293–00111295), 3♀ (00111033, 00111296– 00111297) (CNC). New Mexico: San Juan Co.: 10 mi E of Blanco, 36.724036 N 107.643786 W, 14 Jun 1992, W.F. Chamberlain, 58 (00118257–00118260, 00118379) (TAMU). Oregon: Baker Co.: 2 mi NE of Rt 26 on Rt 245, Unity Lake State Park, 44.486966 N 118.191736 W, 27 Jun 1988, G.M. Stonedahl, Artemisia tridentata , 38 (00107868, 00108059– 00108060), 11♀ (00108061–00108071) (AMNH). Unity, 44.43756 N 118.191676 W, 13 Jul 1991, W.F. Chamberlain, 18 (00118261) (TAMU). Deschutes Co.: 1 mi S of Millican, 43.864716 N 120.918896 W, 1335 m, 21 Jun 1979, R.T. Schuh, Artemisia tridentata ssp. tridentata , 28 (00108056–00108057) (AMNH). 1 mi S of Millican on Pine Mountain Road, 43.864716 N 120.918896 W, 1335 m, 21 Jun 1979, G.M. Stonedahl, G. Cooper, M.D. Schwartz, Artemisia sp. , 18 (00107864) (AMNH); 21 Jun 1979, G.M. Stonedahl, Artemisia tridentata ssp. tridentata , 18 (00108054), 1♀ (00108055) (AMNH). 2.6 mi S of Millican, T20S R15E S10, 43.841516 N 120.918896 W, 1405 m, 21 Jun 1979, M.D. Schwartz, Artemisia tridentata , 28 (00111261–00111262) (CNC). Redmond, 44.272786 N 121.172786 W, 25 Jun 1939, Schuh and Gray, 1♀ (00107998) (AMNH). Harney Co.: Blitzen River, 42.614446 N 119.068066 W, 06 Jul 1906, unknown, 2♀ (00119274–00119275) (UCB), 18 (00134514) (USNM). Fish Lake, Steens Mountains, 42.737626 N 118.64136 W, 2286 m, 22 Jun 1951 – 26 Jun 1951, Borys Malkin, 18 (00242624), 1♀ (00242625) (CAS). Frenchglen, 42.826946 N 118.915556 W, 1372 m, 21 Jun 1951, Borys Malkin, 18 (00242626), 1♀ (00242627) (CAS). Narrows, 43.278846 N 118.962076 W, 1253 m, 01 Jul 1906, unknown, 18 (00134516) (USNM). Warm Springs V[alley], 43.276826 N 119.249126 W, 1253 m, 07 Jul 1906, unknown, 18 (00134515) (USNM). Klamath Co.: 4 mi NW of Worden on road to Keno, 42.090036 N 121.902276 W, 17 Jul 1985, G.M. Stonedahl and J.D. McIver, Artemisia tridentata , 1♀ (00108072) (AMNH). Swan Lake, 42.228616 N 121.598896 W, 12 Jul 1953, J. Schuh, 18 (00118065) (ORSU). Lake Co.: 3 km N of Fort Rock State Park, 43.398866 N 121.068056 W, 1326 m, Jun 1982 – Jul 1982, Neil Cobb, Artemisia tridentata Nutt. (Asteraceae) , 18 (NAUF). 5.7 mi E of Jct. Rts 395–140, Warner Mountain Pass, 42.23446 N 120.291116 W, 1765 m, 03 Jul 1979, R.T. Schuh and B. Massie, Artemisia tridentata , 1♀ (00108058) (AMNH). 11 mi NW of Silver Lake on Hwy 31, 43.247056 N 121.157126 W, 1372 m, 21 Jun 1979, G.M. Stonedahl, Artemisia tridentata , 18 (00107863) (AMNH); 25 Jun 1979, G. Stonedahl, Artemisia tridentata , 58 (00108047–00108051), 2♀ (00108052–00108053) (AMNH); 25 Jun 1979, M.D. Schwartz, Artemisia tridentata ssp. tridentata , 28 (00111256–00111257), 3♀ (00111258–00111260) (CNC). 28 mi SE of Jct. of Rts 97 and 31, 43.299356 N 121.180696 W, 1497 m, 25 Jul 1979, M.D. Schwartz, Artemisia tridentata ssp. tridentata , 118 (00110989, 00111028, 00111244–00111252), 3♀ (00111253– 00111255) (CNC). 28 mi SE of La Pine on Rt 31, 43.37666 N 121.209666 W, 1535 m, 25 Jun 1979, R.T. Schuh, Artemisia tridentata , 98 (00108034–00108042), 4♀ (00108043–00108046) (AMNH). Abert Lake, 42.570246 N 120.187676 W, 02 Jul 1935, Joe Schuh, 18 (00107996) (AMNH). Big Squaw Flat, 18 mi SE of Bly, 42.397786 N 121.040566 W, 25 Jul 1962, J.D. Vertress, 1♀ (00108073) (AMNH). Lakeview, 42.188896 N 120.344726 W, 23 Jun 1933, G.P. Engelhardt, 18 (00117807), 1♀ (00117808) (KU). Paisley, 42.693886 N 120.546116 W, 1332 m, 28 Jun 1951 – 29 Jun 1951, Borys Malkin, 18 (00134533) (USNM). Silver Lake, 43.128066 N 121.0456 W, 1510 m, 15 Jun 1938, Schuh and Gray, 18 (00118064) (ORSU); 16 Jun 1938, Gray and Schuh, 18 (00107997) (AMNH). Summer Lake, 42.827866 N 120.795596 W, 19 Jun 1952, Vincent Roth, 88 (00118055–00118062), 1♀ (00118063) (ORSU). Malheur Co.: 30 mi E of Rome, 42.837646 N 117.034526 W, 26 Jun 1952, R. Lauderdale, 18 (00118067) (ORSU). Succor Creek Canyon, 43.59226 N 117.059826 W, 726 m, 15 Jun 1951 – 18 Jun 1951, Borys Malkin, 18 (00242623) (CAS). Umatilla Co.: 2 mi E of Hermiston, 45.840556 N 119.246766 W, 12 Jun 1963, Charles W. Baker, 18 (00118066) (ORSU). Utah: Cache Co.: 0.4 mi off Rt 89 along Franklin Basin Rd, 41.935556 N 111.572226 W, 2004 m, 17 Jul 2001 – 18 Jul 2001, T.J. Henry and A.G. Wheeler, Jr., Artemisia tridentata , 18 (00122137), 1♀ (00122138) (USNM). 0.4 mi off Rt 89 along Franklin Basin Rd, 41.935556 N 111.572226 W, 2004 m, 18 Jul 2001, T.J. Henry and A.G. Wheeler, Jr., Artemisia tridentata , 18 (00122158), 2♀ (00122159–00122160) (USNM). 5 mi off Rt 89 along Franklin Basin Rd, 41.991666 N 111.591946 W, 2256 m, 18 Jul 2001, T.J. Henry and A.G. Wheeler, Jr., Artemisia tridentata , 4♀ (00122161–00122164) (USNM). 14 mi S on FS Rd 055 off Rt 89, USU Forest, T13N R4E Sec15, 41.546586 N 111.749446 W, 2591 m, 25 Jul 1981, M.D. Schwartz, Artemisia tridentata , 18 (00111298), 2♀ (00111299–00111300) (CNC). 14 mi S on FS Rd 055 off of Rt 89, USU For, T13N R4E Sec15, 41.86886 N 111.48446 W, 2591 m, 25 Jul 1981, M.D. Schwartz, Artemisia tridentata , 1♀ (00111034) (CNC). Logan Canyon, 41.74026 N 111.793836 W, 24 Jul 1948, G.G. Fleener, Castilleja sp. (Scrophulariaceae) , 1♀ (00134526) (USNM). Logan Canyon site, 41.740276 N 111.793056 W, 1981 m, 15 Jul 1958, G.F. Knowlton, 38 (00122165– 00122167) (USNM). Carbon Co.: Minnie Maud Creek, West Tavaputs Plateau, 51.8 mi SW of Myton, T12S R12E, 39.774826 N 110.509596 W, 1981 m, 09 Jul 1982, M.D. Schwartz, Artemisia tridentata , 28 (00111307–00111308) (CNC). Duchesne Co.: 23.7 mi S of Myton, Well’s Draw, T10S R15E, 39.851816 N 110.061116 W, 1829 m, M.D. Schwartz, Artemisia tridentata , 1♀ (00111303) (CNC). Emery Co.: 13.2 mi NW of junction of Hwy 10 on Hwy 31, 39.466856 N 110.857336 W, 2179 m, 17 Jul 1980, G.M. Stonedahl, Artemisia sp. , 88 (00107945– 00107952), 3♀ (00107953–00107955) (AMNH). Grand Co.: South Beaver Mesa, La Sal Mountains , 38.63756 N 109.101116 W, 04 Jul 1980, J.T. and D.A. Polhemus, 18 (00116753) (JTP). Juab Co.: 7 mi W of Eureka, 39.945376 N 112.226126 W, 1841 m, 26 Jun 1966, W.M.T., Artemisia tridentata , 18 (00134506) (USNM). Fernow Valley, Juab County, 39.519726 N 111.939726 W, 06 Jun 1940, R.W. Fautin, 1♀ (00134524) (USNM). Millard Co.: Scipio, 39.2456 N 112.103336 W, 29 Jun 1965, H.H. Knight, 18 (00134508) (USNM). Rich Co.: Garden City, 41.889176 N 111.3856 W, 06 Jul 1968, G.F. Knowlton and H.H. Knight, 1♀ (00134529) (USNM). Salt Lake Co.: Salt Lake City, 40.760836 N 111.890286 W, 27 Jul 1953, R.R. Dreisbach, 18 (00127323) (UMMC). San Juan Co.: 20 mi SE of Blanding, 37.43486 N 109.38846 W, 1571 m, 20 Jun 1999, W.F. Chamberlain, 1♀ (00245352) (TAMU). Abajo Mountains, 0.2 mi SE Devils Canyon Campground, 38.151286 N 109.838516 W, 2103 m, 12 Jun 1982, M.D. Schwartz, Artemisia tridentata , 18 (00111304), 1♀ (00111305) (CNC). Brushy Basin Rim Rd, SW of Blanding, county rd # 227, 0.5 E of milepost 116, T37S R12E, 37.566246 N 109.55346 W, 1737 m, 12 Jun 1982, M.D. Schwartz, Artemisia tridentata , 28 (00111023, 00111306), 2♀ (00111029, 00111309) (CNC). Grand Flat near Collins Canyon, 37.421676 N 110.160566 W, 1707 m, 01 Jun 1982, D.A. and J.T. Polhemus, 48 (00116702, 00116750–00116752) (JTP). Sevier Co.: 2.3 mi N of I-70 on road to Kanosh, 38.606386 N 112.452066 W, 2128 m, 16 Jul 1980, R.T. Schuh and G.M. Stonedahl, Artemisia tridentata , 1♀ (00107988) (AMNH); 16 Jul 1980, G.M. Stonedahl, Artemisia sp. , 48 (00107865, 00107983–00107985), 2♀ (00107986, 00107987) (AMNH). Clear Creek Narrows Summit on Rt 4, 38.581566 N 112.256786 W, 2243 m, 15 Jul 1980, G.M. Stonedahl, Artemisia sp. , 2♀ (00107989– 00107990) (AMNH). Dog Spring Rd at Rt 25, 38.505696 N 111.795236 W, 2718 m, 16 Jul 1980, R.T. Schuh and G.M. Stonedahl, Artemisia tridentata , 68 (00107971–00107976), 1♀ (00107977) (AMNH). Jct. Mytoge Mountain Rd and Hwy 25, 38.513566 N 111.773166 W, 2697 m, 16 Jul 1980, G.M. Stonedahl, Artemisia sp. , 28 (00107978–00107979), 3♀ (00107980–00107982) (AMNH). Salt Gulch, 12 mi N of Hwy 24 on Hwy 72, 38.512926 N 111.524736 W, 2457 m, 17 Jul 1980, G.M. Stonedahl, Artemisia sp. , 1♀ (00107991) (AMNH). Tooele Co.: Benmore, environs, 40.01256 N 112.420836 W, C. Jorgensen, 1♀ (00070014) (USNM); 19 Jun 1954, G.L. Nielson, 1♀ (00134527) (USNM). Uintah Co.: 5–10 mi SW of Bonanza, T10S R24E Sec 29 (G2), 39.91616 N 109.24066 W, 1524 m, 08 Jun 1982, M.D. Schwartz, Artemisia tridentata , 118 (00110994, 00111310–00111319), 7♀ (00111320–00111326) (CNC). 5–10 mi SW of Bonanza, T11S R24E Sec 6, Asphalt Wash, 39.8866 N 109.27216 W, 1615 m, 09 Jun 1981, M.D. Schwartz, Artemisia tridentata , 28 (00111327–00111328), 2♀ (00111329–00111330) (CNC). Blue Mountain Flat and Cliff Ridge, T5S R25E, 40.360836 N 109.131396 W, 2438 m, 08 Jul 1982, M.D. Schwartz, Artemisia tridentata , 18 (00111301), 1♀ (00111302) (CNC). Bonanza, White River Shale Project Trailers, T 9S R24E S23, 39.979516 N 109.179576 W, 1524 m, 01 Jun 1981, M.D. Schwartz, Artemisia tridentata , 28 (00110988, 00111331–00111332), 3♀ (00111333–00111335) (CNC). Utah Co.: Cedar Valley Environs, 40.336846 N 112.103816 W, 1583 m, C. Jorgensen, 28 (00070012, 00134534) (USNM). Mount Nebo, 39.823346 N 111.744466 W, 12 Jul 1942, G.F. Knowlton, 18 (00134537) (USNM). Spanish Fork Canyon, 40.062836 N 111.570696 W, 17 Jul 1954, G.F. Knowlton, 18 (00134535) (USNM). Wasatch Co.: 30 mi SE of Kamas on Rt 35, Uinta National Forest, Wolf Creek Campground, T4S R10W Sec 7, 40.481626 N 111.030376 W, 2743 m, 15 Aug 1986, M.D. Schwartz and G.M. Stonedahl, Artemisia tridentata , 128 (00107869, 00107956–00107966), 4♀ (00107967–00107970) (AMNH). Deer Creek Reservoir, 40.40956 N 111.53036 W, 1677 m, C. Jorgensen, 18 (00070013) (USNM). Heber City, 40.506946 N 111.41256 W, 05 Jul 1922, E.P. Van Duzee, Artemisia sp. , paratypes of Junior synonym ( S. uniformis ), 11♀ (00242596–00242606) (CAS), paratype of Junior synonym ( S. uniformis ), 18 (00134510) (USNM). Washington: Benton Co.: Hanford Site ALE, 117 gate to 1200 Ft Rd, 46.488836 N 119.653336 W, 24 May 1995, R.S. Zack, Artemisia tridentata , 1♀ (00121913) (UID), 28 (00121908– 00121909), 3♀ (00121910–00121912) (WSU). Hanford Site ALE, Rattlesnake Ridge, spring, 46.583896 N 119.386676 W, 23 May 1995, R.S. Zack, 48 (00121902–00121905) (WSU). Hanford Site ALE, radio telescope, 46.583896 N 119.386676 W, 13 Jun 1995, R.S. Zack, 28 (00121906–00121907) (WSU). Yakima Co.: Toppenish, 46.37756 N 120.30756 W, 15 May 1926, E.W. Davis, 28 (00134521–00134522) (USNM). Wyoming: Albany Co.: The Summit, 41.231666 N 105.440556 W, 2591 m, 10 Aug 1950, R.R. Dreisbach, 18 (00127324) (UMMC). Big Horn Co.: Big Horn Mountains, 44.754996 N 107.782396 W, 1981 m, 17 Aug 1927, H.H. Knight, 18 (00134520) (USNM). Carbon Co.: 7 mi SW of Encampment, 41.137676 N 106.884516 W, 2682 m, 05 Aug 1967, F., P. and B. Rindge, 18 (00107928) (AMNH). Saratoga, 41.4556 N 106.805836 W, 29 Jul 1931, H.H. Knight, 18 (00134505) (USNM). Lincoln Co.: Alpine, 43.160126 N 111.023286 W, 1731 m, 19 Jul 1936, G.F. Knowlton, 1♀ (00134536) (USNM). Salt River Pass, 15 mi S of Afton on Rt 89, 42.542346 N 110.894316 W, 2326 m, 21 Jul 1981, M.D. Schwartz, Pinus contorta Dougl. ex Loud. (Pinaceae) , 18 (00111025), Artemisia tridentata , 28 (00111024–00111285), 7♀ (00111286–00111292), (CNC). Park Co.: National Park [Yellowstone], 44.766676 N 110.233336 W, 01 Aug, unknown, 1♀ (00134512), (USNM). Yellowstone National Park, 44.766676 N 110.233336 W, 12 Aug 1907, W. Robinson, 18 (00134509), (USNM). Sublette Co.: 24 mi W of Big Piney, 42.537366 N 110.58556 W, 2560 m, 01 Aug 1928, F., P. and B. Rindge, 18 (00107927), (AMNH). 25 mi SE of Jackson, Hobart River, 43.224436 N 110.410956 W, 27 Jun 1966, W. Gagne and J. Haddock, Artemisia tridentata , 28 (00119270–00119271), (UCB). Green River Lake[s Campground], Wind River Mountains, 43.311696 N 109.868736 W, 2591 m, 01 Aug 1935 – 08 Aug 1935, H. Ruckes, 18 (00134507), (USNM). Sweetwater Co.: 10 mi E of Rock Springs, 41.58756 N 109.008936 W, 04 Jul 1968, H.H. Knight, 18 (00134525), (USNM). 30 mi E of Rock Springs near Point of Rocks, 41.58756 N 108.622366 W, 04 Jul 1968, H.H. Knight, 18 (00134523), (USNM). Teton Co.: Jackson, 43.486 N 110.761676 W, 2134 m, 15 Aug 1961, J.E.R. Stainer, 1♀ (00111069), (CNC). Teton Pass, 43.49756 N 110.954446 W, 06 Aug 1972, L.A. Kelton, Artemisia sp. , 78 (00110987, 00111049–00111054), 14♀ (00111055–00111068) (CNC).