Pygatyphella karimui, Ballantyne & Lambkin, 2009

Ballantyne, Lesley A. & Lambkin, Christine, 2009, Systematics of Indo-Pacific fireflies with a redefinition of Australasian Atyphella Olliff, Madagascan Photuroluciola Pic, and description of seven new genera from the Luciolinae (Coleoptera: Lampyridae), Zootaxa 1997, pp. 1-188 : 86-87

Pygatyphella karimui sp. n.

( Figs 279–281, 284, 289, 290, 298, 299, 304–306)

Holotype. Male. PAPUA NEW GUINEA: Chimbu Pr., 6.30S, 144.50E, Karimui 1080 m, 14.vii.1963, JS ( BPBM).

Paratypes (6). Same data as holotype, 4 males, 2 females ( BPBM) .

Diagnosis. One of the obsoleta ‘complex’ distinguished by the strongly arched and swollen V7, the broad apically truncate MPP, and the very narrow apices of the LL of the aedeagus.

Male. 8.4–9.6 mm long; 2.8–3.3 mm wide; W/L 0.3. Colour ( Fig. 279): Dorsal surface light brown, mostly semitransparent with fat body showing through the cuticle, and brown markings in median area of pronotum (small, paired, and faint), a narrow median dark band on the MS which reaches anterior margin, narrowly dark along the basal ¼ of suture from posterior margin of MS, small brown area at base of elytra between bases of interstitial lines 1 and 2, and two small preapical brown spots, one at posterior end of interstitial line 1 and the other at outer margin; head antennae and palpi very dark; ventral aspect of thorax yellowish except for brown metasternum; legs yellow with brown tarsi; basal ventrites yellowish, V5 dark brown with anterolateral pale patches; LO yellowish, posterior area of V7 yellowish with underlying muscle impressions clearly visible; basal tergites brown, T7 and 8 yellow. Pronotum ( Fig. 284): 1.6–1.8 mm long; 2.9–3.3 mm wide; W/L 1.8; lateral margins with rounded (3/5) or angulate (1/5) convergence, and indentations in lateral margins in 5/5; posterolateral corners angulate, <90° and inclined obliquely to the median line. Elytron ( Fig. 279): parallel-sided; 6.8–7.8 mm long; with 2 well–defined interstitial lines. Head: can be partially retracted into prothoracic cavity, narrowly visible from above in resting position; GHW 1.9–2.0 mm; SIW 0.15mm; SIW/GHW 0.08; ASD<ASW (sockets very close). Abdomen, ventrites ( Figs 304–306): LO retracted into anterior 1/3 of V7 where it may not reach sides, not reaching posterior margin, posterior margin either entire or slightly and narrowly produced in 1/5; V7 arched and swollen; posterolateral corners rounded angulate; muscle impressions extend almost to tip of MPP. MPP moderately broad, apex squarely truncate; dorsal surface with a wide median ridge ( Fig. 306, arrowed). Tergites: T8 ( Fig. 306) very well sclerotised, about as long as wide, thickened lateral margins converge gently posteriorly; anterolateral prolongations elongate, narrow, expanded vertically, more widely so in median area; ventral surface with a slightly developed long median trough with fine lateral margins. Aedeagus ( Figs 289, 290, 298, 299): L/W>3/1; LL/ML moderate; lateral margins of LL converge posteriorly and LL narrow at apices; LL not divergent along dorsal length except near apices, slightly shorter than ML; outer preapical margin of LL slightly produced and rounded, inner apical margin produced, short and narrow; base of LL asymmetrically produced to left; ML lacking teeth at about level of ejaculatory orifice, symmetrical, not curved left horizontally; dorsolateral margins of ML behind apex slightly produced and pointed (may be visible from beneath as tooth like projections ( Figs 289, 290, 298, 299).

Female. 8.8–9.6 mm long. Coloured as for male ( Figs 280, 281) with these exceptions: pronotum lacking any median darker markings (irregularities in colour attributed to underlying muscles), MN very pale in ½, MS with wider median dark band in 1/2, very pale brown along the basal ¼ of suture from posterior margin of MS, small brown area at base of elytra between bases of interstitial lines 1 and 2 in 1/2; V6 with yellowish LO, V7, 8 brownish yellow, semitransparent; posterior margin of V7 broadly emarginated, posterolateral corners rounded, anterolateral areas slightly depressed; median posterior margin of V8 narrowly emarginate, with a shallow slightly depressed groove along its length in 1/2; lateral margins of T8 converge posteriorly, posterior margin straight.

Larva. Not associated.

Etymology. The specific name, the type locality, is considered as a noun in apposition.