Perrona loetschi, Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 109

Perrona loetschi nov. sp.

Figs 32A 1 –A 3 , B 1 –B 2 , C 1 –C 3 , D 1 –D 2 , 5 , 7

Pleurotoma Jouanneti [sic] Des Moul.— Hörnes 1854: 346 (partim), pl. 38, figs 4a–b [non Tomellana jouannetii ( Des Moulins, 1842) ].

Pleurotoma (Clavatula) descendens Hilb. — Hoernes & Auinger 1891 (partim): 355, pl. 48, figs 7–8 [non Perrona descendens ( Hilber, 1879) ].

Type material. Holotype: NHMW 1949 /0005/0102a, SL: 30.1 mm, MD: 13.5 mm, Grund ( Austria), illustrated in Hoernes & Auinger (1891, pl. 48, fig. 8), Figs 32A 1 –A 3 . Paratypes: NHMW 1949 /0005/0102b, SL: 28.4 mm , MD: 12.1 mm, Grund ( Austria) , illustrated in Hoernes & Auinger (1891, pl. 48, figs 7a–b), figs 32B 1 –B 2; NHMW 1855 /0045/0752, SL: 28.0 mm , MD: 12.1 mm, Grund ( Austria) , illustrated in Hörnes (1854, pl. 38, figs 4a–b), figs 32D 1 –D 2; NHMW 2021 /0153/0001, SL: 31.3 mm , MD: 13.6 mm, Grund ( Austria) , figs 32C 1 – C 3; NHMW 2021 /0153/0002, 31 spec., Grund ( Austria) ; NHMW 1855 /0045/0752, 15 spec., Grund ( Austria) ; NHMW1860 /0050/0059, 2 spec., Neuruppersdorf ( Austria) , SMF 351872 /5, Grund ( Austria) .

Type locality. Grund ( Austria); North Alpine-Carpathian Foreland Basin .

Type stratum. Silty sand of the Grund Formation.

Age. Middle Miocene, early Badenian (early Langhian).

Etymology. In honor of Bernd Lötsch, Director General of the Natural History Museum in Vienna from 1994 to 2009.

Diagnosis. Perrona species of medium size, very solid, broad fusiform, with gradate spire, subcylindrical whorls with strongly swollen, weakly tuberculose subsutural collar. Absence of well-defined suprasutural cord. Much reduced spiral sculpture and relatively short siphonal canal.

Description. Shell medium-sized, very solid, broad fusiform with markedly gradate spire; apical angle 38–45°. Protoconch not preserved. Teleoconch of nine whorls. Early teleoconch whorls weakly concave with two narrow, adsutural spiral cords separated by weakly concave central part with close-set opisthocline, comma-shaped axial riblets. Axial riblets fading out on fourth whorl. Later whorls subcylindrical with prominent, sharply-delimited, swollen, rounded subsutural collar bearing poorly defined tubercles, weakly convex below, suprasutural cord very weak, almost entirely covered by succeeding whorl. Delicate spiral cords on spire whorls (rarely preserved in available material). Suture narrowly incised, linear. Last whorl ~65% of total height, subcylindrical. Subsutural collar very broad and swollen, rounded, weakly beaded. Subsutural ramp very narrow, weakly concave. Shoulder rounded, weakly convex below, weakly angled at moderately constricted and concave base. Siphonal fasciole moderately swollen with prominent growth increments. Sculpture extremely weak, fine indistinct cords over base and fasciole. Aperture moderately wide, pyriform. Outer lip not thickened, smooth within; anal sinus moderately deep, moderately narrow, asymmetrically U-shaped, with apex mid-ramp. Siphonal canal moderately long, moderately wide, only slightly deflected to the left with shallowly notched tip. Columella excavated in upper third, straight below, weakly twisted at fasciole, smooth. Columellar and parietal callus thick, sharply delimited, forming broad rim and narrow pseudumbilical chink.

Discussion. This species was confused by Hörnes (1854) with Tomellana jouannetii ( Des Moulins, 1842) , from the Burdigalian of Mérignac in France, and by Hoernes & Auinger (1891) with Perrona descendens ( Hilber, 1879) from the Badenian of the Paratethys Sea. The separation from Tomellana jouannetii is obvious based on the gradate spire, the irregularly beaded subsutural collar (see Peyrot 1931, pl. 8, figs 43, 61, 62, 66, 71) and the early teleoconch sculpture (see discussion on Tomellana ). The separation from Perrona descendens is less obvious at first sight. Nevertheless, Perrona loetschi differs in its more solid, broader shell, shorter spire, the broader ‘shelf’ of the subsutural collar, the lower penultimate whorl and higher last whorl, and the much reduced spiral sculpture. The broad outline and the absence of distinct spiral cords on the entire teleoconch allow also a separation from Perrona vindobonensis (Quenstedt, 1884) and P. ilonae nov. nom. Perrona louisae ( Hoernes & Auinger, 1891) has even broader whorls and differs in its pointed beads.

Paleoenvironment. At the locality Grund fossiliferous channel fills, which formed in middle to outer neritic environments bear allochthonous assemblages uniting coastal-mudflat faunas with inner neritic ones ( Zuschin et al. 2005; Roetzel 2009). Preservation and color of the shells, suggest, that Perrona loetschi belonged to the allochthonous shallow water fauna and not to the autochthonous deep-water fauna.

Distribution in Central Paratethys. Karpatian (early Miocene): North Alpine-Carpathian Foreland Basin: Neuruppersdorf ( Austria) (hoc opus); Badenian (middle Miocene): North Alpine-Carpathian Foreland Basin: Grund ( Austria) ( Hoernes & Auinger 1891).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana).

FIGURE 32 A1–A3. Perrona loetschi nov. sp., holotype, NHMW 1949/0005/0102a, Grund (Austria). B1–B2. Perrona loetschi nov. sp., paralectotype, NHMW 1949/0005/0102b, Grund (Austria). C1–C3. Perrona loetschi nov. sp., paralectotype, NHMW 2021/0153/0001, Grund (Austria). D1–D2. Perrona loetschi nov. sp., paralectotype, NHMW 1855/0045/0752, Grund (Austria). E1–E3. Perrona louisae (Hoernes & Auinger, 1891), lectotype, NHMW 1861/0050/0162a, Weinsteig (Austria). F1–F2. Perrona louisae (Hoernes & Auinger, 1891), paralectotype, NHMW1861/0050/0162b, Weinsteig (Austria).