Perrona ursulae ( Hoernes & Auinger, 1891 )

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 132-133

Perrona ursulae ( Hoernes & Auinger, 1891)

Figs 40D 1 –D 3 , E 1 –E 3 , F 1 –F 3 , 4W , 5 , 7

Clavatula Ursulae —Hoernes 1891: 131 [nomen nudum].

* Pleurotoma (Clavatula) Ursulae nov. form.— Hoernes & Auinger 1891: 358, pl. 48, figs 19–21.

Pleurotoma (Clavatula) ursulae R. H. et Au.— Zalányi 1913: 545.

Clavatula (Perrona) jouanneti [sic] Desm.— Florei 1961: 680, pl. 6, fig. 40 [non Tomellana jouanetii ( Des Moulins, 1842) ].

Perrona (Perrona) ursulae ( Hoernes et Auinger, 1891) —Hinculov in Iliescu et al. 1968: 148, pl. 37, figs 6–7.

Perrona (Perrona) ursulae (Hoernes & Auinger) — Nicorici 1972: 69.

Perrona (Perrona) vindobonensis (Partsch) — Atanacković 1985: 167, pl. 38, figs 3–4 [non Perrona vindobonensis ( Hoernes & Auinger, 1891) ].

non Clavatula ursulae R. Hörnes und Auinger, 1891 — Moisescu 1955a: 168, pl. 13, figs 18–19 [= Perrona oliviae ( Hoernes &

Auinger, 1891)].

Type material. Lectotype designated herein: NHMW 1949 /0005/0107a, Buituri ( Romania), SL: 33.4 mm, MD: 12.3 mm, illustrated in Hoernes &Auinger (1891, pl. 48, fig. 20), figs 40F 1 –F 3 . Paralectotype: NHMW 1949 /0005/0107b, Buituri ( Romania), SL: 22.8 mm , MD: 8.3 mm, figs 40E 1 –E 3; NHMW 1846 /0001/1294, Buituri ( Romania), SL: 32.7 mm , MD: 10.9 mm, figs 40D 1 –D 3, 4W ; NHMW 1862 /0001/0552, 20 spec., Buituri ( Romania) ; NHMW 1949 /0005/0107c, Buituri ( Romania), SL: 23.8 mm , MD: 8.8 mm, illustrated in Hoernes & Auinger (1891, pl. 48, fig. 19).

Additional studied material. NHMW 1861 /0033/0043, 19 spec., Buituri ( Romania) .

Revised description. Shell medium-sized, solid, moderately broad fusiform with high spire; apical angle 30– 33°. Protoconch not preserved. Teleoconch of at least 12 whorls. Early teleoconch whorls with narrow indistinctly beaded subsutural cord, weakly concave mid-portion with strongly opisthocline axial riblets, with swollen abapical ends forming suprasutural row of beads. On seventh whorl subsutural cord strengthens to form prominent, sharply delimited, subsutural collar, riblets at mid-whorl disappear replaced by fine spiral cords. Suprasutural cord narrow, partly covered by succeeding whorl. On ninth whorl subsutural collar strong, somewhat swollen in the lower half; concave mid-whorl portion narrows, cords weaken. Suprasutural cord broadens, flattened, weakly bisected by faint spiral groove. Suture narrowly impressed, linear. Last whorl ~55% of total height. Subsutural collar strong, rounded, very prominent. Subsutural ramp narrow. Shoulder high-placed, weakly angular, mid-whorl portion inflated. Base weakly angled, delimited by weak cord. Base strongly constricted, siphonal fasciole poorly developed. Strong spiral cords over base of alternating strength, strengthened perifasciolar cord; cords of alternate strength over fasciole. Aperture moderately wide, pyriform. Outer lip not thickened, with about eight coarse lirae deep within; anal sinus moderately deep, narrow, asymmetrically U-shaped, with apex mid-ramp; siphonal canal moderately long and wide, deflected to the left with shallowly notched tip. Columella excavated in upper third, weakly twisted at fasciole, smooth. Columellar and parietal callus thin, adherent, well delimited from base, forming broad rim.

Discussion. Perrona ursulae ( Hoernes & Auinger, 1891) is characterized by its very strongly delimited and roundly swollen subsutural collar that is somewhat further thickened in the abapical half giving the impression of overhanging the narrow concave subsutural ramp and the very strongly developed cords over the base of alternating strength. It differs from Perrona lydiae ( Hoernes & Auinger, 1891) in the well-defined subsutural collar and the presence of lirae inside the aperture. Perrona oliviae ( Hoernes & Auinger, 1891) is more slender fusiform and also lacks lirae. Tomellana jouannetii ( Des Moulins, 1842) , as defined by Peyrot (1931), develops a similar subsutural collar, but has a much higher base, a shorter spire and lacks comma-shaped riblets on early teleoconch whorls, placing it in the genus Tomellana rather than Perrona which has comma-shaped axial riblets on the neanic whorls.

Paleoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin: Hrvaćani ( Bosnia and Herzegovina) ( Atanacković 1985); Făget Basin: Buituri ( Romania) ( Boettger 1906; Zalányi 1913); Șimleu Basin: Tusa, Vînători ( Romania) ( Nicorici 1972); Mahida Basin: Valea Bela Reca, Valea Calvei ( Romania) (Hinculov in Iliescu et al. 1968); Zorlenţul Mare Basin ( Romania) ( Florei 1961).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 7. Apertures of Paratethyan Clavatulidae (O. = Olegia, ‘C.’ = ‘Clavatula’, N. = Neoperrona, P. = Perrona, Pu. = Pusionella, Sc. = Scaevatula, S. = Striopusionella, T. = Tomellana).

FIGURE 40 A1–A3. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309a, Pöls (Austria). B1–B3. Perrona styriaca (Hilber, 1879), paralectotype, NHMW 1846/0037/0309b, Pöls (Austria). C1–C3. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). D1–D3, Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). E1–E3. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0107b, Buituri (Romania). F1–F3. Perrona ursulae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0107a, Buituri (Romania).