Dactyladenia floretii Breteler, 2000

Breteler, F. J., 2000, Novitates Gabonenses 38. Two new species of Dactyladenia (Chrysobalanaceae) from Gabon, Adansonia (3) 22 (1), pp. 117-122 : 117-118

Dactyladenia floretii Breteler , sp. nov.

Dactyladeniae jongkindii Breteler maxime simile, sed bracteolis minoribus 3 -palmatilobatis ad invicem 5- palmatilobatis et pedicello infimo multo brevi differt; a speciebus aliis congeneribus stipulis saccatis differt.

TYPUS. — Breteler, Dibata, Mayombo & Moussavou 14490, Gabon, near Gamba, fl., fr. July (holo-, WAG; iso-, G, LBV, MO, P).

Shrub, lianescent shrub, or liana. Main stem up to 8 cm diam. Branches hispid mixed with short erect hairs or with short erect hairs only, rarely with hispid indumentum only. Stipules saccate, shortly stipitate, slightly curved or straight, ovoidellipsoid in outline, (5-)8-20 × (2-) 6-8 mm, glabrous or with some arachnoid hairs sometimes mixed with some hispid hairs, margin distinctly and densely pubescent. Leaves: petiole ± terete, 2- 4(-5) mm long, hairy as branchlets; lamina coriaceous to papyraceous, oblong-elliptic, ovate-elliptic to oblong-obovate, sometimes narrowly so, (6-)10-25(-27) × (2-)4-7(-8) cm, 2-4(-5) times as long as wide, narrowly and usually deeply cordate at base, acute to shortly and acutely acuminate at apex, the acumen up to 0.5(1.5) cm long; glabrous above usually with some erect hairs or an arachnoid remnant on the impressed midrib, glabrous or hispid beneath especially so on midrib and main laterals, glabrescent or not, the (8-)10- 14(15) pairs of main lateral nerves and tertiairy venation plane or slightly prominent above, usually prominent beneath; glands ± flat, present only beneath, at some distance alongside the midrib and some more dispersed ones, more densely concentrated on the leaf apex and near base. Inflorescence a simple, ± erect raceme, up to 7 cm long, usually axillary, sometimes terminal, few- to many-flowered, puberulous; peduncle with a few empty bracts; bracts ovate-triangular, soon reflexed, 4-5 mm long, arachnoid-hairy and with puberulous midrib beneath, glabrous above, the margin with 1-2 glands on each side, puberulous; lower pedicel (2) 4-7 mm long, puberulous. Bracteoles ± insert- ed at the top of the lower pedicel, deeply 3- palmatilobate, 1-2 mm long, puberulous, lobes gland-tipped. Upper pedicel 1.5-3 mm long, puberulous; receptacle tube (11-) 16-20 mm long, gibbous at base, puberulous, sometimes almost glabrous outside, inside with some hispid hairs at mouth. Sepals ovate-oblong to elliptic, 5-8 × 2.5- 4 mm, with acute to emarginate apex, tomentose inside as well as on the outside parts covered in bud, puberulous to almost glabrous outside; the outer sepals sometimes with a few flat glands. Petals white, early caducous, thin, oblong-elliptic, 10-15 × 5-8 mm, with a short, glabrous claw, up to 1 mm long. Stamens 20-28, 4- 4.5 cm long, glabrous, filaments united for ca. 2/3 of their length; anthers ca. 0.6 mm long. Style slightly longer than the stamens, glabrous, with a small stigma; ovary long-velutinous at apex. Fruit ovoid, narrowly so or not, rounded at both ends, 3.5-6 × 2-2.5 cm, velutinous and with some longer hispid hairs. — Figs. 1 , 2.

PARATYPES. — GABON: Breteler 9457, Rabi- Kounga, ster. Mar. ( WAG) ; Breteler & Jongkind 10219, eod. loc., ster. Oct. ( WAG) ; Breteler , Dibata , Mayombo & Moussavou 14435, Gamba, fl. July ( LBV, P, WAG) ; J. J. de Wilde & van der Maesen 10907, Rabi-Kounga, fr. Jan. ( WAG) ; Floret & Louis 1419, Mviadi , NW of Kango, fl. Sept. ( P, WAG) ; Louis 2212, Mayumba, fl. Oct. ( WAG) ; Wieringa 1130, Gamba, ster. June ( WAG) ; Wieringa & van de Poll 1320, eod. loc., fl. July ( WAG) .

NOTE. — This species is named after J.-J. FLORET, the first who recognized that the material he collected with A.M. LOUIS represented a new species.

Figures

Fig. 1. — Dactyladenia floretii: A, leafy branch with erect inflorescences; B, inflorescence; C, stipule; D, flower; E, bracteole; F, fruit. (A, C-E, Breteler et al. 14435; B, F, Breteler et al. 14490). Drawing by H. de VRIES.

Fig. 2. — Distribution of Dactyladenia floretii (•) and D. jongkindii (*).