Bolbelasmus (Bolbelasmus) makrisi Miessen, 2011

Sommer, David, Hillert, Oliver, Hrůzová, Lucie & Král, David, 2021, Bolbelasmus (Bolbelasmus) zagrosensis (Coleoptera: Scarabaeoidea: Bolboceratidae), a new species from Iran, along with an updated key to the western Palaearctic species of the subgenus, Zootaxa 4920 (3), pp. 380-394 : 385

Bolbelasmus (Bolbelasmus) makrisi Miessen, 2011

( Figs. 1B , 2B , 3B , 4C , 5B , 9 )

Bolbelasmus (Bolbelasmus) makrisi Miessen, 2011: 111 , figs. on pp. 116, 118–119 [not numbered] (type locality: “ Kourion , Lemesos, alt.: 50 m, Cyprus ”).

Type material examined. Cyprus: Paratypes, 1 ♁ 1 ♀ ( GMCL), “Rizoelia nat. for. Park | Larnaka—CYPRUS || leg.: E. Kakouris | collection: G. Miessen || Bolbelasmus makrisi nov. sp. | G. Miessen det. 2011 | PARATYPUS [printed, red label]” .

Additional material examined. Cyprus: 1 ♁ 1 ♀ ( OHCB), Akamas NP, 5 km N of Lara, 13.–20.iii.2015, O. Hillert lgt.; 1 ♁ ( OHCB), Akamas NP, Lara , 13.–20.iii.2015, O. Hillert lgt.; 1 ♁ ( OHCB), Miliou, N of Paphos, 13.–20.iii.2015, O. Hillert lgt.; 1 ♁ ( OHCB), Miliou, 34°56’22.23”N, 32°27’35.90”E, 7.vi.2014, vivant dans piscine [alive in pool], G. Miessen lgt.; 1 ♁ ( OHCB), Miliou, 34°56’21.66”N, 32°27’35.59”E, 7.vi.2014, attire par lampe UV [attracted by UV light], G. Miessen lgt.; 16 ♁♁ 15 ♀♀ ( OHCB) ( Figs. 1B , 2B , 3B , 4C , 5B ), 2 ♁♁ 2 ♀♀ ( NMPC), 5 km NE of Pegeia , Paphos env., 13.–20.iii.2015, O. Hillert lgt .

Distribution. Endemic to Cyprus ( Nikolajev et al. 2016) (see map, Fig. 9 ).

Figures

FIGURE 1A–D. Habitus of males in dorsal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale.

FIGURE 2A–D. Pronotum of males in frontal view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale.

FIGURE 3A–D. Habitus of males in lateral view. A, holotype of Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895) from Greece (Rhodes); D, B. nireus from Turkey. Not to scale.

FIGURE 4A–D. Pronotum of males in lateral view. A, Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species, from Fars province, Iran; B, B. zagrosensis Sommer, Hillert, Hrůzová & Král, new species, from Lorestan province, Iran; C, B. makrisi Miessen, 2011; D, B. nireus (Reitter, 1895) from Greece, Rhodes. Not to scale.

FIGURE 5A–C. Parameres in dorsal view. A, Bolbelasmus zagrosensis Sommer, Hillert, Hrůzová & Král, new species; B, B. makrisi Miessen, 2011; C, B. nireus (Reitter, 1895). Not to scale.

FIGURE 9. Sketch map with marked distribution of Bolbelasmus makrisi Miessen, 2011 (green), B. nireus (Reitter, 1895) (yellow) and B. zagrosensis Sommer, Hillert, Hrůzová & Král, new species (red). Squares represents the type locality of each species.