Cyclocephala rufa Endrödi, 1967

Neita-Moreno, Jhon C., 2021, A review of the black species of Cyclocephala Dejean (Coleoptera: Scarabaeidae Dynastinae) from Colombia, Zootaxa 5026 (1), pp. 1-58 : 39-44

publication ID	https://doi.org/ 10.11646/zootaxa.5026.1.1
publication LSID	lsid:zoobank.org:pub:07E0C922-6B0F-4916-85C2-AD95146A8F1E
DOI	https://doi.org/10.5281/zenodo.10240024
persistent identifier	https://treatment.plazi.org/id/03932D70-B605-C35C-FF7B-0A14A526DBCE
treatment provided by	Plazi
scientific name	Cyclocephala rufa Endrödi, 1967
status

Treatment

( Figs. 3D View FIGURE 3 ; 15I View FIGURE 15 ; 16I View FIGURE 16 ; 17I View FIGURE 17 ; 18K View FIGURE 18 ; 19G View FIGURE 19 ; 25A View FIGURE 25 ; 29E–F View FIGURE 29 ; 40 View FIGURE 40 )

Cyclocephala carinatipennis Martínez & Morón, 1984: 48–52 . NEW SYNONYMY. The character states described for both species are conspecific, and so C. carinatipennis is placed in junior synonymy.

Diagnosis. In C. rufa and C. arhuacana , the surface of the pygidium has sparse punctures ( Fig. 25A–B View FIGURE 25 ), but in C. dalensi , C. kaszabi , and C. ligyrina , the surface of the pygidium has dense punctures ( Fig. 24F–H View FIGURE 24 ). Cyclocephala rufa can be distinguished by the larger punctures on the pygidium (small in C. arhuacana ). The parameres ( Fig. 29E, F View FIGURE 29 ) are also different from those of C. arhuacana ( Fig. 30A–B View FIGURE 30 ).

Redescription. Male ( Fig. 3D View FIGURE 3 ). Length 22.0– 22.7 mm; width 10.1–10.8 mm. Color dark reddish brown to black. Head: Surface of frons and clypeus with punctures moderately dense, deep, and moderately large except those on occiput and apex of clypeus that are smaller and slightly denser. Clypeus with anterior angles broadly rounded, apex broadly truncate, weakly emarginate. Mandibles apically rounded, internal face with a groove; labium densely setose, paraglossa undeveloped, apex slightly notched; maxilla with galea developed, with 6 teeth. Interocular width equals 3.1–3.3 transverse eye diameters. Antenna with 10 antennomeres, club subequal in length to antennomeres 2–7. Pronotum: Surface with punctures as on frons. Base lacking marginal bead. Elytra: Surface with distinct rows of punctures: punctures (and those in intervals) ocellate, moderate to moderately large, moderately dense. Pygidium: Surface with dense punctures, moderate in size, most with a minute, tawny seta in unworn specimens. In lateral view, surface regularly convex ( Fig. 25A View FIGURE 25 ). Legs: Protibia tridentate, basal tooth small and directed “forward” and slightly removed from others ( Fig. 18K View FIGURE 18 ). Protarsus enlarged, tarsomeres 2–4 successively gradually larger, 5th large, only weakly curved ( Fig. 17I View FIGURE 17 ), with large basal tooth; median claw large, strongly curved ( Fig. 16I View FIGURE 16 ), broadly split into a wide ramus and a slender ramus ( Fig. 15I View FIGURE 15 ); metatibia on external face with 2 rows of setae, second row crosses the external face of metatibia ( Fig. 19G View FIGURE 19 ). Venter: Prosternal process long, stout, apex obliquely flattened into a wide, transverse oval with anterior 4/5 raised as convex “button”. Genitalia: Parameres as in Fig. 29E–F View FIGURE 29 .

Female. According to Martínez & Morón (1984), the female is similar to the male except for the following characters: stronger punctures on the dorsal region of the head; epipleuron (ventral view) gradually tapering and then abruptly constricted at metacoxa; in dorsal view, epipleuron gradually expands to level of third sternite; tooth on ventral surface of epipleuron absent in lateral view. Protibia with teeth sharper than in males. Sternite with stronger punctures than in males.

Distribution. Cyclocephala rufa is known from Colombia, Venezuela, and Ecuador ( Endrödi 1967; Martínez & Morón 1984; Ratcliffe et al. 2020).

Life history. Adults are active during the night and attracted to lights.

Locality records ( Fig. 40 View FIGURE 40 ). Two specimens examined from CTNI and ICN‒MHN: “ Colombia, César, / Jagua de Ibirico, Alto de las / Flores. Marzo 10-1996 / H. Restrepo, Leg / Alt. 1204. // Cyclocephala rufa Endrödi, 1967 , Det. J. C. Neita, 2007” [♂ -ICN-028934]. “ Colombia, César, / Jagua de Ibirico, Alto de las / Flores. Marzo 10-1996 / H. Restrepo, Leg / Alt. 1204. // Cyclocephala rufa Endrödi, 1967 , Det. J. C. Neita, 2007” [♂ -CTNI].