Ocypus (Pseudocypus), Mulsant & Rey, 1876

Published, First, 2007, Contributions to the knowledge of the “ Staphylinus-complex ” (Coleoptera: Staphylinidae: Staphylinini) of China. Part XX. The genus Ocypus Leach, 1819, subgenus Pseudocypus Mulsant & Rey, 1876. Section 1, Zootaxa 1421, pp. 1-72 : 7-10

Key to Chinese species of subgenus Pseudocypus of the O. semenowi lineage

1 Pronotal hypomeron microsetose, number of microsetae varying from a few to quite numerous ( Figs. 213–215) ................................................................................................................................................... 2

- Pronotal hypomeron glabrous, lacking microsetae................................................................................. 20

2 Legs uniformly rufobrunneous ................................................................................................................. 3

- Legs not uniformly rufobrunneous, at least partially darkened, or entirely dark, piceous to black ......... 7

3 Visible abdominal tergites 4 and 5 with patch of not dense, golden-yellowish tomentose pubescence covering almost entire tergite. Aedoeagus as in Figs. 97–102. Length 11.0–14.0 mm. (Shaanxi: Qinling Shan) .................................................................................................................. O. neocles sp. nov. (pars)

- No patches of yellowish tomentose pubescence on abdominal tergites. Aedoeagi different ................... 4

4 Male antenna short, segment 4 about as long as wide, outer segments wider than long ( Fig. 216). Underside of paramere without blackish sensory peg setae (Fig. 163). Aedoeagus as in Figs. 161–163. Length 11.0–14.0 mm. (Gansu, Qinghai, Shaanxi, northern Sichuan) ............... O. semenowi Reitter, 1887 (pars)

- Male antenna relatively long, segment 4 distinctly longer than wide (ratios 1.27–1.40) ( Fig. 217), outer segments as long as wide, or longer as wide. Underside of paramere with blackish sensory peg setae (Figs. 169, 175). Aedoeagi different ......................................................................................................... 5

5 Apical portion of paramere narrow, with subacute to acute apex (Figs. 168, 169). Aedoeagus as in Figs. 167–169. Length 14.0–18.0 mm. (Sichuan) ................................................................. O. glabrio sp. nov.

- Apical portion of paramere wide, with apex arcuate to subtruncate (Figs. 175, 178, 181)...................... 6

6 Apex of median lobe narrowly arcuate (Figs. 178, 179). Apical portion of paramere markedly asymmetrical (Figs. 178, 180, 181). Postgena sparsely punctate and pubescent ( Fig. 212). Length 16.0–18.0 mm. (Southern Sichuan) .......................................................................................................... O. ballio sp. nov.

- Apex of median lobe subtruncate (Fig. 174). Apical portion of paramere slightly asymmetrical (Figs. 173, 175). Postgena densely punctate and pubescent ( Fig. 211). Length 13.0–15.0 mm. (Central Sichuan) ....................................................................................................................... O. teuthras sp. nov.

7 Visible abdominal tergites 4 and 5 each with patch of yellowish tomentose pubescence in middle........ 8

- No patches of yellowish tomentose pubescence on abdominal tergites ................................................. 11

8 Apex of both median lobe and paramere very wide, widely arcuate; paramere large and wide, covering most of median lobe ( Figs. 200–202). Length 16.0–17.0 mm. (Sichuan: Emei Shan, Erlang Shan, Gongga Shan) .................................................................................................................. O. pelias sp. nov.

- Apex of both median lobe and paramere moderately wide, variably arcuate, obtusely rounded, subacute or subtruncate, that of median lobe sometimes emarginate in middle (Figs. 98, 106, 112); paramere moderately large and wide, never covering most of median lobe (Figs. 97, 105, 111) ................................... 9

9 Apex of median lobe narrowly emarginate in middle (Figs. 111, 112). Aedoeagus as in Figs. 111–114. Small species. Length 10.0–13.0 mm. (Shaanxi: Qinling Shan) .................................. O. rhinton sp. nov.

- Apex of median lobe not emarginate in middle (Figs. 98, 106). Aedoeagi different ............................. 10

10 Apex of paramere almost reaching apex of median lobe (Fig. 97); apex of median lobe obtuse to subtruncate (Fig. 98). Sternite 9 of male genital segment subtruncate apically (Fig. 95). Golden-yellowish tomentose pubscence on visible abdominal tergites 4 and 5 not dense, covering almost entire tergite. Length 11.0–14.0 mm. (Shaanxi: Qinling Shan) ............................................... O. neocles sp. nov. (pars)

- Apex of paramere distinctly not reaching apex of median lobe (Fig. 105); apex of median lobe narrowly arcuate to subacute (Figs. 105, 106). Sternite 9 of male genital segment emarginate apically (Fig. 103). Golden-yellowish tomentose pubescence on visible abdominal tergites 4 and 5 dense, limited to middle of tergite. Length 12.0–15.0 mm. (Hubei: Da Shennongjia range) ..................... O. puer (Smetana, 2005)

11 Paramere markedly elongate, shaped variably ( Figs. 129, 136, 142, 153, 157)..................................... 12

- Paramere moderately elongate, shaped variably ( Figs. 117, 184, 190) .................................................. 18

12 Paramere with subacute apex markedly exceeding apex of median lobe (Fig. 157). Underside of paramere without black sensory peg setae (Fig. 158). Length 15.0 mm. (Gansu: Wenxian) ..................... ....................................................................................................................................... O. abaris sp. nov.

- Paramere with apex of different shape, no more than moderately exceeding apex of median lobe ( Figs. 129, 136, 142, 153). Underside of paramere with black sensory peg setae ( Figs. 132, 138, 145, 154). 13

13 Apical portion of paramere conspicuously asymmetrical, of characteristic shape ( Figs. 129, 131, 132). Length 15.0–18.0 mm. (Sichuan: Maokian, Wenchuan) .................................................... O. itys sp. nov.

- Apical portion of paramere of different shape ( Figs. 138, 145, 154) ..................................................... 14

14 Apical portion of median lobe narrow, with subacute apex not reaching apex of median lobe ( Figs. 149, 195) ......................................................................................................................................................... 17

- Apical portion of median lobe wider, with obtusely arcuate apex about reaching apex of paramere ( Figs. 136, 142, 153) ......................................................................................................................................... 15

15 Apical portion of paramere markedly curved mediad ( Figs. 136, 142), sensory peg setae on underside of paramere extended markedly posteriad along lateral margins ( Figs. 138, 145) ..................................... 16

- Apical portion of paramere not appreciably curved mediad (Fig. 153), sensory peg setae on underside of paramere forming two short lateral groups connected anteriorly (Fig. 154) Length 16.0 mm. (Sichuan: Lixian) ........................................................................................................................... O. gorgias sp. nov.

16 Apical portion of paramere narrow, conspicusously curved mediad, apex of paramere not reaching apex of median lobe ( Fig. 142); right margin of paramere (paramere removed from median lobe and viewed underside up) appearing obtusely angulate ( Figs. 144, 145). Length 15.0–18.0 mm. (Sichuan: Xuecheng) ................................................................................................................................ O. xerxes sp. nov.

- Apical portion of paramere moderately wide, less conspicuously curved mediad, apex of paramere about reaching apex of median lobe ( Fig. 136); right margin of paramere (paramere removed from median lobe and viewed underside up) appearing almost evenly arcuate ( Figs. 137, 138). Length 16.0– 18.0 mm. (Sichuan: Lixian) ......................................................................................... O. elpenor sp. nov.

17 Apical portion of paramere narrow, with apex irregularly, narrowly arcuate ( Figs. 195, 197, 198). Length 20.0 mm. (Sichuan: Heishui Shan) .................................................................... O. orodes sp. nov.

- Apical portion of paramere wide, with apex widely rounded ( Figs. 149, 150). Length 16.0 mm. (Sichuan: Lixian) ..................................................................................................... O. pammenes sp. nov.

18 Apex of median lobe markedly exceeding apex of paramere ( Fig. 190). Aedoeagus as in Figs. 190, 191. Length 19.0–20.0 mm. (Sichuan: Wenchuan) .................................................................... O. bion sp. nov.

- Apex of median lobe about reaching apex of paramere ( Figs. 117, 184). Aedoeagi different............... 19

19 Apex of median lobe subtruncate with slight protuberance near apex of paramere ( Fig. 118). Aedoeagus moderately large, as in Figs. 117–120. Sternite 9 of male genital segment arcuate apically ( Fig. 115). Smaller species. Length 16.0–20.0 mm. (Shaanxi: Daba Shan) ...................................... O. dolon sp. nov.

- Apex of median lobe obtuse, with characteristic formation on face adjacent to paramere on left side below apex ( Fig. 185). Aedoeagus large, as in Figs. 184–187. Sternite of male genital segment emarginate apically (Fig. 182). Larger species. Length 21.0–23.0 mm. (Sichuan: Jiuding Shan) .......................... ................................................................................................................................. O. palamedes sp. nov.

20 Legs uniformly rufotestaceous to rufobrunneous ................................................................................... 21

- Legs not uniformly rufotestaceous to rufobrunneous, at least partially darkened, or entirely dark, piceous to black....................................................................................................................................... 30

21 Visible abdominal tergites 4 and 5 each with patch of yellowish tomentose pubescence (mostly not conspicuous). Aedoeagus as in Figs. 3–7. Length 13.0–16.0 mm. (Sichuan) ........................ O. zetes sp. nov.

- No patches of yellowish tomentose pubescence on abdominal tergites ................................................. 22

22 Aedoeagus heavily pigmented, appearing black, shaped as in Figs. 59, 60. Surface of head and pronotum with slight purplish tint. Length 13.0–16.0 mm. (Sichuan) ................................. O. scaevola sp. nov.

- Aedoeagus not heavily pigmented, appearing brownish, shaped differently. Surface of head and pronotum without appreciable purplish tint ..................................................................................................... 23

23 Male antenna short, segment 4 about as long as wide, outer segments wider than long ( Fig. 216). Aedoeagus as in Figs. 161–163. Length 11.0–14.0 mm. (Gansu, Qinghai, Shaanxi, northern Sichuan) ............. ......................................................................................................................... O. semenowi Reitter (pars)

- Male antenna relatively long, segment 4 distinctly longer than wide (ratios 1.15–1.30) ( Fig. 217). Aedoeagi different................................................................................................................................... 24

24 Underside of paramere without blackish sensory peg setae ( Figs. 22, 79)............................................. 25

- Underside of paramere with blackish sensory peg setae ( Figs.14, 36, 43, 50, 126)............................... 26

25 Paramere short, apex situated considerably below apex of median lobe ( Fig. 77). Aedoeagus as in Figs. 77–79, in general appearing short. Interspaces between punctures on head and pronotum without microsculpture. Length 13.0–16.0 mm. (Yunnan: Dali) ................................................... O. hyas sp. nov.

- Paramere elongate, apex situated near apex of median lobe ( Fig. 17). Aedoeagus as in Figs. 17–22, in general appearing elongate. Interspaces between punctures on head and pronotum with microsculpture. Length 15.0–16.0 mm. (Sichuan) .............................................................................. O. sarpedon sp. nov.

26 Apex of median lobe wide, widely arcuate ( Fig. 12). Aedoeagus as in Figs. 11–14). Length 12.0–16.0 mm. (Sichuan) ............................................................................................................... O. laelaps sp. nov.

- Apex of median lobe narrow, subacute to narrowly arcuate ( Figs. 34, 40, 47, 123).............................. 27

27 Sensory peg setae on underside of paramere not numerous (about 10), forming a small apical field not extended posteriad along lateral margins of paramere ( Fig. 126). Aedoeagus as in Figs. 123–126. Length 15.0–18.0 mm. (Shaanxi: Daba Shan) ...................................................................... O. menander sp. nov.

- Sensory peg setae on underside of paramere numerous to very numerous, in general forming two lateral groups connected anteriorly, or an extensive apical field extended posteriad along lateral margins of paramere ( Figs. 36, 42, 49) ..................................................................................................................... 28

28 Aedoeagus elongate, paramere long and narrow, with conspicuous longitudinal crest on face away from median lobe, apex of paramere about reaching apex of median lobe ( Fig 34). Sternite 9 and tergite 10 of male genital segment elongate ( Figs. 32, 33). Length 15.0–17.0 mm. (Sichuan) O. quiris sp. nov. (pars)

- Aedoeagus less elongate, paramere moderately long and wide, without conspicuous longitudinal crest on face away from median lobe, apex of paramere not reaching apex of median lobe ( Figs. 40, 47). Sternite 9 and tergite 10 of male genital segment not elongate ( Figs. 38, 39, 45, 46) .................................. 29

29 Apex of paramere situated considerably below apex of median lobe; sensory peg setae on underside of paramere forming two lateral groups connected anteriorly ( Figs. 40, 43). Aedoeagus as in Figs. 40–43. Length 13.0–17.0 mm. (Sichuan) .................................................................. O. sericeomicans Bernhauer

- Apex of paramere situated close to apex of median lobe; sensory peg setae forming an extensive apical field extended posteriad along lateral margins of paramere ( Figs. 47, 49). Aedoeagus as in Figs. 47–50. Length 14.0–18.0 mm. (Sichuan: Wa-wu Shan) ........................................................... O. imurai Smetana

30 Aedoeagus heavily pigmented, appearing black, shaped as in Figs. 71–74. Length 15.0–16.0 mm. (Sichuan) ...................................................................................................................... O. calamis sp. nov.

- Aedoeagus not heavily pigmented, appearing brownish, shaped differently ......................................... 31

31 Interspaces between punctures on head and pronotum without appreciable microsculpture, surface therefore relatively shiny. Aedoeagus as in Figs. 83–86. Small species. Length 10.0–12.0 mm. (Xinjiang) ................................................................................................................................ O. dryas sp. nov.

- Interspaces between punctures on head and pronotum with microsculpture, surface therefore more or less dull. Aedoeagi different. Larger species (except for O. zeuxis ), length exceeding 13.0 mm .......... 32

32 Sensory peg setae on underside of paramere pale, not pigmented ( Fig. 55). Aedoeagus as in Figs. 54, 55. Length 13.0–16.0 mm. (Gansu) .................................................................................... O. sadales sp. nov.

- Sensory peg setae on underside of paramere pigmented, appearing black ( Figs. 30, 36, 67, 93). Aedoeagi different............................................................................................................................................. 33

33 Paramere moderately elongate, apical portion of median lobe markedly asymmetrical, with acute apex (Fig. 90). Small, slender species. Length 11.0–13.0 mm. (Hubei: Da Shennongjia range)........................ ........................................................................................................................................ O. zeuxis sp. nov .

- Paramere markedly elongate, apical portion of median lobe of different shape ( Figs. 26, 34, 64). Larger species. Length 15.0–19.0 mm ............................................................................................................... 34

34 Sensory peg setae on underside of paramere very numerous, extended considerably posteriad along left lateral margin ( Fig. 36); paramere on face away from median lobe with conspicuous longitudinal crest ( Fig. 34). Length 15.0–17.0 mm. (Sichuan) ......................................................... O. quiris sp. nov. (pars)

- Sensory peg setae on underside of paramere moderately numerous, no more than moderately extended posteriad along right lateral margin ( Figs. 30, 67); paramere without conspicuous crest on face away from median lobe, but with a fine carina ( Figs. 26, 64).......................................................................... 35

35 Apical portion of paramere constricted into a narrow, acute apex ( Figs. 64, 66, 67). Sternite 9 of male genital segment deeply emarginate apically ( Fig. 62). Aedoeagus as in Figs. 64–67. Length 16.0–19.0 mm. (Sichuan: Sabde) .................................................................................................... O. hecato sp. nov.

- Apical portion of paramere with subacute, triangular apex ( Figs. 26, 29, 30). Sternite 9 of male genital segment only slightly emarginate apically ( Fig. 24). Aedoeagus as in Figs. 26–30. Length 15.0–18.0 mm. (Sichuan: Zhilong) ................................................................................................. O. vindex sp. nov.