Adelomompha Minet & Park, 2020

Park, Kyu-Tek, Koo, Jun-Mo & Minet, Joël, 2020, Review of the Malagasy lecithocerid species described by Pierre Viette and deposited in MNHN (Paris), with new generic combinations and descriptions of a new subfamily and genus of Momphidae (Lepidoptera: Gelechioidea), Zootaxa 4845 (2), pp. 151-190 : 181-182

Adelomompha Minet & Park , gen. nov.

LSID: urn:lsid:zoobank.org:act:8A37619B-8B8A-4E5E-AA44-6114F2DB9335

Type species: Lecithocera andrianella Viette, 1968a: 88 .

Diagnosis. Differs from the other momphid genera in antenna length (exceeding forewing length in both sexes of Adelomompha ), number of forewing M veins (two rather than three), shape of the abdominal spinose areas (wide and entire on the tergal surface while typically divided mesally in Mompha and relatives), shape of the valva in male genitalia (valva entire while longitudinally split in most Momphinae ), and shape of the signum in female genitalia (a small, plare-like structure in Adelomompha ; a pair of signa including, or composed of, sickle-shaped structures in all Momphinae but two Palaeomystella species, in which the corpus bursae lacks signa). It should be noted, however, that the valva has lost its longitudinal cleft in one species of Mompha endemic to the Falkland Islands (Wakeham-Dawson & Koster 2013).

Description. Head: Head capsule evenly convex, with smooth scale vestiture; on vertex, scales fairly broad, with rounded distal edges. Compound eyes relatively small, remote from each other and distinctly notched near the almost adjacent antennal foramina. Ocelli absent.Antenna much longer than forewing and devoid of well developed sensilla ventrally (in both sexes); scape elongate, tapering proximally, without pecten or any projecting hair-like scale; dorsal surface of each flagellomere with two rows of small, narrow scales; in female, proximal region of the flagellum slightly thickened with scales dorsally. Labial palpi large, slender, sickle-shaped, entirely smooth-scaled (while most Momphinae have a projecting vestiture at the apex of segment 2); third segment arched and pointed, almost as long as second. Maxillary palpi folded over base of proboscis; the latter moderately long, scaled for most of its length.

Thorax: Foreleg well developed; coxa long; tibia approximately as long as femur and provided, near its middle, with an epiphysis that does not extend beyond the extremity of the tibia (epiphysis slightly shorter in female); tarsus as long as tibia; male with a long hair pencil that arises near the base of the coxa and is partially hidden between the latter and the mesothorax. Midleg: tibia medium-sized, smooth-scaled, with a pair of unequal spurs (long spur and basitarsus have approximately the same length); tarsus slightly longer than tibia. Hindleg: tibia longer than midtibia, in male with a conspicuous dorsal fringe of very long piliform scales; mesal spurs markedly unequal (long spur about twice as long as short spur), arising slightly before (i.e. proximad of the middle of the tibia; apical spurs less unequal; tarsus slightly longer than tibia. All legs: dorsal scales of the distitarsus overhanging the pretarsus (which is very small). Forewing with only two M veins and longitudinally folded along the upper M vein (which is in a more ventral position than the lower M vein and ends well below the apex); R 1 long, arising approximately from the middle of the upper edge of the discal cell; R 2 arising closer to R 3 than to R 1 but remote from upper angle of discal cell; R 3 free, arising near stem R 4 +5; R 5 running to costa; CuA 1 and CuA 2 separate, the former arising near the lower M vein; CuP long, distally present as a tubular vein; anal vein short, its extremity remote from that of CuP. Female wing coupling: retinaculum of subcostal type, that is composed of hooked scales attached to the base of vein Sc; frenulum bisetose. Male frenulum a strong bristle. Hind wing lanceolate; vein Sc+ R 1 long, relatively straight; Rs running to costa, ending well before apex; M 1 and M 2 short-stalked; stem M 1+2 separate from Rs but closer to it than to the lower angle of the discal cell, from which arise M 3 and CuA 1; CuA 1 and CuA 2 separate, fairly remote; lower edge of the discal cell distinctly arched proximad of CuA 2 and somewhat parallel to the anal margin; vein CuP long. Metascutum smooth-scaled posteriorly, without piliform scales. Anterior edge of metascutellum angled on meson (cephalad) .

Abdomen: First tergum with a well developed neotergite; terga II-VII anteriorly provided with wide, undivided areas covered in spiniform scales. Sternum II with a pair of anterolateral processes (with heavily pigmented mesal edges in the male) and well indicated venulae that do not reach the apodemes; the latter moderately long in the female but nearly absent in the male. Segment VIII of male: tergum reduced; sternum large, produced anteriorly on meson and with its anterolateral angles extended, forming narrow arms (thus resembling the eighth sternum of Palaeomystella rosaemariae Moreira & Becker, 2014 : Fig. 5C in Luz et al., 2014).

Male genitalia ( Figs. 22G, H ): aedeagus long, its outer wall desclerotized, except for a heavily pigmented dorso-longitudinal band and two posterolateral sclerites, which may represent remnants of a juxta; one strong clawlike cornutus; vinculum composed of thin arms and not produced into a saccus; valva broad, entire, rounded distally, provided with a very large ring-shaped structure; transtilla absent; tegumen broad in lateral view, separated from uncus by a deep groove; gnathos absent; uncus elongate, strongly bent proximally and minutely pointed at its posterior extremity.

Female genitalia ( Figs. 22 I–K): corpus bursae more or less ovoid, with a small, toothed, plate-like signum provided with at least one lateral point; ductus bursae slender, longer than corpus; ostium bursae remote from segment VIII; the latter setose, without free ventral lobes; apophyses anteriores fairly long, although much shorter than the apophyses posteriores; papillae anales forming an elongate ovipositor.

Figures

FIGURE 5. Torodora ankasokella (Viette, 1968), comb. nov.: A, holotype; B, labels; C, male genitalia, prep. J. Minet no. 1662; D, aedeagus; E, abdomen. Scale bar for the genitalia and aedeagus: 0.5 mm.

FIGURE 22. Adelomompha andrianella (Viette, 1968), comb. nov.: A, holotype of andrianella Viette; B and B', ditto, ventral surface of fore- and hind wings; C, ditto, labels; D, holotype of ranavaloella Viette; E, ditto, labels; F, ditto, lateral view of head; G, male genitalia, prep. J. Minet no. 1664; H, ditto, aedeagus; I, female genitalia (posterior region), prep. P. Viette no. 4754; J, ditto, bursa copulatrix; K, ditto, close-up of signum; L, abdomen of the male. Scale bar: 0.5 mm.