Hiya

Kessler, Michael, Smith, Alan R., Øllgaard, Benjamin, Matos, Fernando B. & Moran, Robbin C., 2023, Prodromus of a fern flora of Bolivia. XLII. Update I., Phytotaxa 630 (3), pp. 183-210 : 185

Hiya

Schwartsburd et al. (2017) treated 12 species of Hypolepis for Bolivia, including Hypolepis nigrescens Hook. More recently, Shang et al. (2018) conducted a phylogenetic analysis of Hypolepis and related genera and found that three species placed in Hypolepis ( H. brooksiae , H. nigrescens , and H. scabristipes ) are more closely related to the genera Blotiella , Paesia , and Histiopteris than to the remainder of Hypolepis . They therefore created the new genus Hiya to accommodate these three species. Hiya is characterized by indeterminate, scandent leaves exhibiting rhythmic growth, provided with recurved black-tipped prickles, and stipule-like pinnules that protect the emerging crozier and young pinna apices, rachis-costa architecture where the adaxial sulcus is confluent with the next lower order, and a base chromosome number of x = 29 ( Shang et al. 2018). The name for what we called Hypolepis nigrescens thus is: