Agnotecous Saussure, 1878

Desutter-Grandcolas, Laure, Anso, Jérémy & Jourdan, Hervé, 2016, Crickets of New Caledonia (Insecta, Orthoptera, Grylloidea): a key to genera, with diagnoses of extant genera and descriptions of new taxa, Zoosystema 38 (4), pp. 405-452 : 444

Genus Agnotecous Saussure, 1878

Agnotecous Saussure, 1878: 546 .

TYPE SPECIES. — Agnotecous tapinopus Saussure, 1878 by original monotypy.

ILLUSTRATIONS. — Habitus: Fig. 15D . Male genitalia: Desutter- Grandcolas & Robillard 2006: figs 4-6. Calling songs: Robillard et al. 2010: figs 3, 7.

DIAGNOSIS. — After Otte et al. (1987), Desutter-Grandcolas (1997c) and Desutter-Grandcolas & Robillard (2006). Medium to large species with brown coloration, without HWs and with short HWs in both males and females. Head flat dorsally. Fastigium very wide, 1.5 to 2 × as large as scape. Ocelli set as a wide and flat triangle; median ocellus vertical and subapical on fastigium. Eyes not prominent. Scapes small, slightly longer than wide. Joint 5 of maxillary palpi short, regularly widened distad, truncate obliquely at apex. Pronotum distinctively long and wide; DD somewhat wider than long, narrowed anteriorly. TI with an oval outer tympanum and a slit-like inner tympanum; with three apical spurs.TII with four apical spurs. TIII with four subapical spurs on each side; inner subapical spurs longer and strongly curved; with three inner and three outer apical spurs, median spur the longest on each side; median outer spur twice as long as other outer spurs.TIII serrulated over its whole length. Basitarsomeres III with two rows of dorsal spines; inner row with one or few spines. Cerci moderately long.

Male. Metanotum without glandular structures. FWs short, not reaching abdomen mid length, truncate dorsally; lateral field very high, longer than dorsal field, and partly covering body dorsum. Venation: dorsal field with complete stridulum, with one or no harp vein, and a reduced mirror, lost in apical reticulation; lateral field with numerous parallel veins. Subgenital plate long and rounded; apex narrow and acute; lateral margin inflated, probably glandular. Male genitalia: Pseudepiphallic sclerite very elongate, tubular; pseudepiphallic apex most often elongate as a prominent median process, and with two distal lobes; rami short, extending pseudepiphallic sclerite; pseudepiphallic parameres small and little differentiate; ectophallic fold simple and very short; ectophallic apodemes long, thin and almost parallel; dorsal cavity wide, low and distinctly plicated transversally; endophallic sclerite located at level of ectophallic arc; endophallic apodeme crest-like on dorsal cavity.

Female. FWs short, not reaching tergite II distal margin; most often not overlapping; dorsal and lateral fields with strong, longitudinal parallel veins, and variable transverse veinlets. Subgenital plate small, transverse. Ovipositor straight. Female genitalia with a very small copulatory papilla; apex sclerotized, distinctly separate from half-ringed basal sclerite.

CALLING SONGS. — Long echemes emitted with a dominant frequencies of 15 to 18 kHz, carried by the second harmonic ( Anso et al. 2016b).

DISTRIBUTION. — Endemic to New Caledonia. Mostly forest-dwelling species known throughout the whole Grande Terre.

HABITAT. — Forest-dwelling species which usually hide in leaf litter, and are active at night either in the leaf litter or perched on understorey plants.

Figures

FIG. 15. — New Caledonian Euscyrtinae Gorochov,1985 (A), Oecanthinae Blanchard,1845 (B) and Eneopterinae Saussure,1874 (C-F) (Gryllidae):A, Proturana sp., from Sarramea, by day (Photo PG); B, Oecanthus rufescens Serville, 1839, from Col des Roussettes, by night (Photo JA); C, Cardiodactylus novaeguineae (de Haan, 1842), from Lifou (Loyalty Islands), by night (Photo PG); D, Agnotecous sp., from Col de Mouirange, by day (Photo LDG); E, Lebinthus lifouensis Desutter- Grandcolas, 1997, from Lifou, by day (Photo PG); F, Pixibinthus sonicus Anso & Robillard, 2016, from Col de la Madeleine, by day (Photo JA). Scale bars: 2 mm.