Ficus dulciaria Dugand (1942: 40)

Pederneiras, Leandro Cardoso, Mansano, Vidal De Freitas & Romaniuc-Neto, Sergio, 2017, Taxonomic synopsis of the Ficus sect. Pharmacosycea (Moraceae) from Colombia, Phytotaxa 313 (1), pp. 1-22 : 7

7. Ficus dulciaria Dugand (1942: 40) .

Type:— COLOMBIA. Putumayo: alta cuenca del Río Putumayo, en el valle de Sibundoy, bosque paramero en el filo de la Cordillera, La Cabaña , 2 January 1941, J. Cuatrecasas 11647 (lectotype designated here: COL3876 !; isolectotypes: COL3877! F! US!) .

= Ficus dulciaria var. antioquensis Dugand (1942: 40) . Type:— COLOMBIA. Antioquia: Municipio de Támesis, Cordillera Occidental , cabeceras del Río Cartama , February1942, J. B. Londoño s. n. (lectotype designated here: COL3879 !; isolectotypes: COL3878 ! US!).

= Ficus huilensis Dugand (1942: 51) . Type:— COLOMBIA. Huila: Cordillera Oriental , vertiente occidental, entre Gabinete y Andalucia, 24 March 1940, J. Cuatrecasas 8588 (holotype: COL!; isotypes: F! US!).

Terrestrial trees, 8–25 m tall. Leaf blade 14–21 cm long, 6.0– 12 cm wide, elliptic or broad elliptic, base cuneate to rounded, apex acute, face adaxial puberulent, face abaxial hispid; secondary veins 20–24 pairs; petiole 3.0–4.0 cm long, glabrous, periderm persistent; stipules caducous, 6.5–8.0 cm long, glabrous. Syconia solitary; peduncle 10–15 mm long, glabrous; lateral bracts present; receptacle 2.7–4.0 cm in diam. when dry, hispid, apex rounded ( Fig. 4A, B ).

Distribution and habitat in Colombia: — Antioquia, Caldas, Cauca, Huila, Quíndio, Putumayo, Risaralda, Tolima, in montane rainforests at elevations of 1800–2900 m ( Fig. 2 ).

Selected specimens: — COLOMBIA. ANTIOQUIA: Belmira, Vereda Playas, bosque acueducto Puerto Pingo, Roldán 3495 (HUA); Envigado, Cerro Astillero, Finca La María, Correa 1286 (COL, HUA, JAUM); Jardín, Carretera Jardín-Riosucio, Ventanas, Churchill 15843 (HUA, NY); Peque, quebrada El Aquilón hacia arriba, Benitez 642 (COL, JAUM); Londoño s.n. (COL); Yarumal, Tuberquia 2341 (HUA).— CALDAS: Manizales, Reserva Torre Cuatro, Alvear-P. 321 (COL).— CAUCA: Parque Nacional Munchique, El Tampo, Barbosa 8557 (COL).— QUINDÍO: Cordoba, Vélez 5649 (COL).— RISARALDA: Pereira, Parque Regional Ucumari, Galeano 272 (COL); Pereira, Parque Regional Ucumarí, El Cedral, Vargas 809 (HUA); Pereira, Parque Ragional Ucumari, entre El Cedral y La Pastora, Bernal 1695 (COL); Pereira, Parque Regional Ucumari, Luteyn 13149 (COL, NY). TOLIMA: Ibague, corregimiento Toche, Agudelo 3454 (COL); 9 Km de Murillo hacia Líbano, Escobar 4181 (HUA).

Notes:— There are two specimens of Ficus dulciaria (COL 3876, COL 3877) and two of Ficus dulciaria var. antioquensis (COL 3878, COL 3879) in Herbario Nacional Colombiano (COL) labelled as type. According to the ICNAFP ( McNeill et al. 2012, Art. 9.1), a holotype of a name of a species is a single specimen, i.e., a single herbarium sheet (Art.8.2) or more than one preparation with the parts clearly labelled as being part of the same specimen (Art.8.3). Dugand (1942) did not labelled these specimens as being part of a single specimen, thus, they are duplicates and lectotypes are needed (Art. 9.1 and 9.2). We choose the specimen COL 3876 as a lectotype for F. dulciaria because this herbarium sheet has syconia; and the specimen COL 3879 as a lectotype for F. dulciaria var. antioquensis because it is the best specimen comprising a stem.

Figures

FIGURE 2. Distribution of Ficus aequatoriales, F. apollinaris, F. carchiana, F. crassiuscula, F. crassivenosa, F. dewolfii and F. dulciaria in Colombia based on herbarium specimens.

FIGURE 4. Twigs with syconia and stipules. A, B. Ficus dulciaria (Barbosa 8557, Correa 1286). C. F. gigantosyce (Garcia-Barriga 15335). D. F. insipida (Zarucchi 5028).